GlyCosmos Portal

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GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	August 18, 2024

Displaying entries **626 - 650** of **1128** in total

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GlyTouCan ID	IUPAC Condensed	Motifs	Subsumption Level ▲	Monoisotopic Mass	Organisms
G67642SS	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex Polysialic acid	Linkage defined saccharide	2587.92	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G67935MX	Gal(b1-3)[Gal(a1-3)Gal(b1-6)]GalNAc(?1-	O-Glycan core 1 fuzzy a-Gal antigen	Linkage defined saccharide	707.25	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G67915MO	GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-4)][Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1681.62	Amniota Archelosauria Archosauria Artiodactyla Aves Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Capra hircus Capra Caprinae Catarrhini Chordata Coelurosauria Coturnix japonica Coturnix Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Lagomorpha Laurasiatheria Leporidae Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neognathae Opisthokonta Oryctolagus cuniculus Oryctolagus Ovis aries Ovis Pecora Perdicinae Phasianidae Phasianinae Primates Rattus Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms unclassified entries unclassified sequences unidentified
G68062BK	Fuc(a1-2)Gal(b1-4)GlcNAc(b1-3)[Neu5Ac(a2-6)]GalNAc(?1-	N-acetyllactosamine O-Glycan core 3 fuzzy Type 2 LN	Linkage defined saccharide	1023.38	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G68253RD	GlcNAc(b1-3)[Fuc(a1-2)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-6)]GalNAc(?1-	Blood group H (type 2) Leb Lewis b Ley O-Glycan core 4 fuzzy	Linkage defined saccharide	1081.42	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Primates Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G68313WI	Neu5Ac(a2-4)Gal(b1-3)[Neu5Ac(a2-6)]GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-4)Gal(b1-3)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan complex Type 1 LN	Linkage defined saccharide	2513.88	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G68668TB	Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan high mannose	Linkage defined saccharide	1558.54	50 kb inversion clade Amniota Artiodactyla Artverviricota Ascomycota Aspergillaceae Aspergillus Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Cavia porcellus Cavia Caviidae Chordata Craniata Deuterostomia Dikarya Dipnotetrapodomorpha Embryophyta Euarchontoglires Eukaryota Eumetazoa Euphyllophyta Eurotiales Eurotiomycetes Eurotiomycetidae Euteleostomi Eutheria Fabaceae Fabales Friend murine leukemia virus Fungi Gammaretrovirus Glires Gnathostomata Gunneridae Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Hystricomorpha Laurasiatheria Magnoliopsida Mammalia Mesangiospermae Metazoa Muridae Murinae Murine leukemia virus Muroidea Mus musculus Mus Mus Myomorpha NPAAA clade Opisthokonta Ortervirales Orthoretrovirinae Papilionoideae Pararnavirae Pecora Pentapetalae Pezizomycotina Phaseoleae Phaseolus vulgaris Phaseolus Primates Retroviridae Revtraviricetes Riboviria Rodentia Ruminantia Sarcopterygii Simiiformes Spermatophyta Streptophyta Streptophytina Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Tracheophyta Vertebrata Viridiplantae Viruses cellular organisms eudicotyledons fabids indigoferoid/millettioid clade leotiomyceta rosids saccharomyceta
G68772UL	GlcNAc6S(b1-6)?-GalNAc-ol		Linkage defined saccharide	506.14	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G68774DG	Gal(b1-3)GlcNAc(b1-2)Man(a1-3)[Gal(b1-3)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan complex Type 1 LN	Linkage defined saccharide	1640.59	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G68983XG	Gal(b1-4)GlcNAc(b1-3)[Gal(b1-4)GlcNAc(b1-6)]Gal(b1-4)Glc(b1-	Paragloboside	Linkage defined saccharide	1072.38	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Primates Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G69044AJ	GalNAc?S(b1-4)GlcNAc(b1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan core basic	Linkage defined saccharide	1542.5	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G69161BS	Neu5Gc(a2-8)Neu5Gc(a2-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)Glc(b1-	Paragloboside	Linkage defined saccharide	1321.43	Amniota Artiodactyla Bilateria Boreoeutheria Bovidae Caprinae Carnivora Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Felidae Feliformia Felinae Felis catus Felis Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Ovis aries Ovis Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G69411IG	Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan core basic N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1275.46	Amniota Amphibia Anura Archelosauria Archosauria Artiodactyla Aves Batrachia Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Lithobates pipiens Lithobates Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neobatrachia Neognathae Opisthokonta Pecora Phasianidae Phasianinae Primates Rana Ranidae Ranoidea Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms
G70046PH	Glc(?1-?)Gal(b1-	Galactosylceramide	Linkage defined saccharide	342.12	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G70081SZ	Neu5Ac(a2-6)Gal(b1-4)[Neu5Ac(a2-6)]GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex Polysialic acid	Linkage defined saccharide	3170.11	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G70073SG	GlcNAc(b1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan core basic	Linkage defined saccharide	1113.41	Acer pseudoplatanus Acer Acereae Achelata Alpharhabdovirinae Amniota Amphibia Amphiesmenoptera Amphipyrinae Anas sp. Anas Anatidae Anatinae Anseriformes Anura Archelosauria Archosauria Arthropoda Artiodactyla Aves Batrachia Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Coelurosauria Coturnix coturnix Coturnix Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Crustacea Decapoda Deuterostomia Dicondylia Dinosauria Dipnotetrapodomorpha Ditrysia Dorylaimia Ecdysozoa Embryophyta Endopterygota Enoplea Euarchontoglires Eucarida Eukaryota Eumalacostraca Eumetazoa Euphyllophyta Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Glossata Gnathostomata Gunneridae Haplorrhini Haploviricotina Heteroneura Hexapoda Hippocastanoideae Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Laurasiatheria Lepidoptera Lithobates pipiens Lithobates Magnoliopsida Malacostraca Mammalia Mandibulata Mesangiospermae Mesocricetus auratus Mesocricetus Metazoa Monjiviricetes Mononegavirales Multicrustacea Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Negarnaviricota Nematoda Neobatrachia Neognathae Neolepidoptera Neoptera Nicotiana alata Nicotiana Nicotianeae Nicotianoideae Noctuidae Noctuoidea Obtectomera Opisthokonta Orthornavirae Palinuridae Palinuroidea Panarthropoda Pancrustacea Panulirus interruptus Panulirus Pecora Pentapetalae Perdicinae Phasianidae Phasianinae Pleocyemata Plusiinae Primates Protostomia Pterygota Ranidae Ranoidea Rhabdoviridae Riboviria Rodentia Ruminantia Sapindaceae Sapindales Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Solanaceae Solanales Spermatophyta Spodoptera frugiperda Spodoptera Streptophyta Streptophytina Teleostomi Tetrapoda Theria Theropoda Tracheophyta Trichinella spiralis Trichinella Trichinellida Trichinellidae Trichoplusia ni Trichoplusia Vertebrata Vesicular stomatitis virus Vesiculovirus Viridiplantae Viruses asterids cellular organisms eudicotyledons lamiids malvids rosids unclassified Anas unclassified Vesiculovirus
G70213NR	Gal(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)Glc(b1-	Paragloboside	Linkage defined saccharide	869.3	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Primates Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G70374BH	GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc-ol	N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1626.61	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Equidae Equus caballus Equus Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Lagomorpha Laurasiatheria Leporidae Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Oryctolagus cuniculus Oryctolagus Pecora Perissodactyla Primates Rodentia Ruminantia Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G70444FA	Gal(b1-3)[Neu5Ac(a2-6)]GalNAc(?1-	O-Glycan core 1 fuzzy	Linkage defined saccharide	674.24	Actinopteri Actinopterygii Amniota Amphibia Anura Artiodactyla Batrachia Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Bufo japonicus Bufo Bufonidae Catarrhini Cercopithecidae Cercopithecinae Cercopithecoidea Chordata Clupeocephala Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dipnotetrapodomorpha Equidae Equus caballus Equus Euarchontoglires Eukaryota Eumetazoa Euteleosteomorpha Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Hyloidea Lagomorpha Laurasiatheria Leporidae Macaca radiata Macaca Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Neobatrachia Neopterygii Opisthokonta Oryctolagus cuniculus Oryctolagus Osteoglossocephalai Pecora Perissodactyla Primates Protacanthopterygii Rana dalmatina Rana Rana Ranidae Ranoidea Rattus norvegicus Rattus Rhinella arenarum Rhinella Rodentia Ruminantia Salmonidae Salmoniformes Salmoninae Salvelinus leucomaenis pluvius Salvelinus leucomaenis Salvelinus Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G70473TT			Linkage defined saccharide	876.13	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G70630DR	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-4)]Man(a1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-	Mannose O-Mannose Polysialic acid Type 1 LN	Linkage defined saccharide	2472.85	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G70868QU	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-6)]Man(a1-3)[Gal(b1-3)Gal(b1-4)GlcNAc(b1-2)[Gal(b1-3)Gal(b1-4)GlcNAc(b1-4)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan complex Polysialic acid	Linkage defined saccharide	3423.21	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G70874WW	GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)Glc(b1-	Paragloboside	Linkage defined saccharide	910.33	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Pecora Primates Rattus Rodentia Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G71105IW	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)[Gal(a1-3)Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)[Gal(a1-3)Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN a-Gal antigen	Linkage defined saccharide	3423.21	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G71125UE	Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)[Gal(a1-3)Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN a-Gal antigen	Linkage defined saccharide	2313.84	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms