GlyCosmos Portal

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GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	August 18, 2024

Displaying entries **876 - 900** of **2574** in total

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GlyTouCan ID	IUPAC Condensed	Motifs	Subsumption Level ▲	Monoisotopic Mass	Organisms
G09287HW	Fuc(a1-2)Gal(b1-3)[Gal(?1-?)GlcNAc(b1-6)]GalNAc(a1-	H antigen (type 3) O-Glycan core 2	Linkage defined saccharide	894.33	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G08520NM	GlcNAc(b1-2)Man(a1-3)[Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan hybrid	Linkage defined saccharide	1437.51	Amniota Amphibia Anura Archelosauria Archosauria Artiodactyla Aves Batrachia Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Lithobates pipiens Lithobates Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Neobatrachia Neognathae Opisthokonta Pecora Phasianidae Phasianinae Primates Rana Ranidae Ranoidea Rattus norvegicus Rattus Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostomi Tetrapoda Theria Theropoda Vertebrata Viruses cellular organisms
G09026AJ		N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2354.86	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Rattus norvegicus Rattus Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G09416DG	Glc(a1-3)Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan core basic	Linkage defined saccharide	1396.49	Amniota Ascomycota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dikarya Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Fungi Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Rodentia Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales Saccharomycetes Saccharomycotina Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms saccharomyceta
G09568JT	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2370.86	50 kb inversion clade Amniota Batoidea Bilateria Boreoeutheria Butea monosperma Butea Catarrhini Chondrichthyes Chordata Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dipnotetrapodomorpha Elasmobranchii Embryophyta Euarchontoglires Eukaryota Eumetazoa Euphyllophyta Euteleostomi Eutheria Fabaceae Fabales Glires Gnathostomata Gunneridae Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Magnoliopsida Mammalia Mesangiospermae Mesocricetus auratus Mesocricetus Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha NPAAA clade Opisthokonta Papilionoideae Pentapetalae Phaseoleae Primates Rodentia Sarcopterygii Simiiformes Spermatophyta Streptophyta Streptophytina Teleostomi Tetrapoda Tetronarce californica Tetronarce Theria Torpedinidae Torpediniformes Tracheophyta Vertebrata Viridiplantae cellular organisms eudicotyledons fabids indigoferoid/millettioid clade rosids
G09420NN	Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1802.64	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Rattus Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G09441VY	GlcNAc(a1-4)Gal(b1-3)[Fuc(a1-2)Gal(b1-4)GlcNAc(b1-6)]GalNAc(a1-	N-acetyllactosamine O-Glycan core 2 Type 2 LN	Linkage defined saccharide	1097.41	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Rattus norvegicus Rattus Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G09517ZR	Neu5Gc(?2-?)Gal(?1-?)GlcNAc(?1-?)[Neu5Gc(?2-?)Gal(?1-?)GlcNAc(?1-?)]Man(a1-3/6)[Gal(?1-?)[Fuc(?1-?)]GlcNAc(?1-?)Man(a1-3/6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	N-Glycan core basic	Linkage defined saccharide	2912.02	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G09552LA	Fuc(a1-2)Gal(b1-3)[Fuc(a1-4)]GlcNAc(b1-3)Gal(b1-4)Glc(b1-	Blood group H (type 1) Lactotriaosylceramide Lc3 Leb Lewis b	Linkage defined saccharide	999.36	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Primates Rattus Rodentia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G09675DY	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc-ol	N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2224.8	Amniota Archelosauria Archosauria Artiodactyla Aves Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Equidae Equus caballus Equus Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Lagomorpha Laurasiatheria Leporidae Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neognathae Opisthokonta Oryctolagus cuniculus Oryctolagus Pecora Perissodactyla Phasianidae Phasianinae Primates Rattus Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms
G09724ZC	Man(a1-3)[Man(a1-3)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan core basic	Linkage defined saccharide	1072.38	Amniota Archelosauria Archosauria Ascomycota Aspergillaceae Aspergillus awamori Aspergillus Aves Bilateria Boreoeutheria Catarrhini Chordata Coelurosauria Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dikarya Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Eurotiales Eurotiomycetes Eurotiomycetidae Euteleostomi Eutheria Fungi Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Neognathae Opisthokonta Pezizomycotina Phasianidae Phasianinae Primates Rattus norvegicus Rattus Rodentia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms leotiomyceta saccharomyceta
G09921WG	Gal(b1-3)GlcNAc(b1-2)Man(a1-3)[Gal(b1-3)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	N-Glycan complex Type 1 LN	Linkage defined saccharide	1786.65	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Rattus norvegicus Rattus Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G10203DW	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(?1-	3-Sialyl-LN (type 2)	Linkage defined saccharide	674.24	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Primates Rattus Rodentia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms unclassified entries unclassified sequences unidentified
G10228OD	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	N-Glycan complex Polysialic acid	Linkage defined saccharide	2733.97	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Cricetidae Cricetinae Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muroidea Myomorpha Opisthokonta Primates Rodentia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G10385EY	Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	I' antigen N-Glycan complex	Linkage defined saccharide	2898.04	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Rattus norvegicus Rattus Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G09730EH	Gal(b1-3)GlcNAc(b1-2)Man(a1-3)[Gal(b1-3)GlcNAc(b1-2)[Gal(b1-3)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan complex Type 1 LN	Linkage defined saccharide	2151.78	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Rattus Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G10179FS	Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc-ol	N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1642.61	Amniota Articulavirales Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Canidae Caniformia Canis lupus familiaris Canis lupus Canis Capra hircus Capra Caprinae Carnivora Catarrhini Cavia porcellus Cavia Caviidae Chordata Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dipnotetrapodomorpha Equidae Equus caballus Equus Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Hystricomorpha Insthoviricetes Lagomorpha Laurasiatheria Leporidae Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Negarnaviricota Opisthokonta Orthomyxoviridae Orthornavirae Oryctolagus cuniculus Oryctolagus Ovis aries Ovis Pecora Perissodactyla Polyploviricotina Primates Rattus Riboviria Rodentia Ruminantia Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms unclassified Orthomyxoviridae unidentified influenza virus
G10013HB	Neu5Gc(?2-?)Gal(?1-3)[GlcNAc(?1-6)]GalNAc(a1-	Tn antigen	Linkage defined saccharide	893.31	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Rattus norvegicus Rattus Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G10219AA	GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-2)[GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan complex	Linkage defined saccharide	1519.57	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Primates Rodentia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G10163TY			Linkage defined saccharide	1109.26	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Cetacea Chondrichthyes Chordata Craniata Deuterostomia Dipnotetrapodomorpha Elasmobranchii Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Pecora Rattus Rodentia Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata Whippomorpha cellular organisms
G10604KJ	Gal(b1-3)[Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-6)]GalNAc(?1-	Lewis x Lex N-acetyllactosamine O-Glycan core 2 fuzzy SSEA-1 Type 2 LN	Linkage defined saccharide	894.33	Ambystoma maculatum Ambystoma Ambystomatidae Amniota Amphibia Artiodactyla Batrachia Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Caudata Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Lagomorpha Laurasiatheria Leporidae Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Oryctolagus cuniculus Oryctolagus Pecora Primates Rodentia Ruminantia Salamandroidea Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G10666VQ	Gal(b1-4)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1964.7	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G10805TH	GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc-ol		Linkage defined saccharide	1318.5	Acer pseudoplatanus Acer Acereae Amniota Anas platyrhynchos Anas Anatidae Anatinae Anseriformes Archelosauria Archosauria Artiodactyla Aves Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Coelurosauria Coturnix japonica Coturnix Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Embryophyta Euarchontoglires Eukaryota Eumetazoa Euphyllophyta Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Gunneridae Haplorrhini Hippocastanoideae Hominidae Homininae Hominoidea Homo sapiens Homo Lagomorpha Laurasiatheria Leporidae Magnoliopsida Mammalia Mesangiospermae Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neognathae Opisthokonta Oryctolagus cuniculus Oryctolagus Pecora Pentapetalae Perdicinae Phasianidae Phasianinae Platanaceae Platanus Primates Proteales Rodentia Ruminantia Sapindaceae Sapindales Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Spermatophyta Streptophyta Streptophytina Teleostomi Tetrapoda Theria Theropoda Tracheophyta Vertebrata Viridiplantae cellular organisms eudicotyledons malvids rosids
G11041DA	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2077.75	Amniota Artiodactyla Bilateria Boreoeutheria Catarrhini Cavia porcellus Cavia Caviidae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Hystricomorpha Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Primates Rattus norvegicus Rattus Rodentia Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G10470SE	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan complex Polysialic acid	Linkage defined saccharide	3390.2	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Mesocricetus auratus Mesocricetus Metazoa Muroidea Myomorpha Opisthokonta Primates Rodentia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms