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GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	August 18, 2024

Displaying entries **1 - 25** of 85 in total

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GlyTouCan ID ▲	IUPAC Condensed	Motifs	Subsumption Level	Monoisotopic Mass	Organisms
G00051MO	Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-	Lewis x Lex N-acetyllactosamine SSEA-1 Type 2 LN	Linkage defined saccharide	529.2	Amniota Bacteria Bifurcata Bilateria Boreoeutheria Campylobacterales Campylobacterota Catarrhini Chordata Colubroidea Craniata Deuterostomia Dipnotetrapodomorpha Elapidae Elapinae Episquamata Epsilonproteobacteria Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Helicobacter pylori 26695 Helicobacter pylori Helicobacter Helicobacteraceae Hominidae Homininae Hominoidea Homo sapiens Homo Lepidosauria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Naja Opisthokonta Primates Rodentia Sarcopterygii Sauria Sauropsida Serpentes Simiiformes Squamata Teleostomi Tetrapoda Theria Toxicofera Unidentata Vertebrata cellular organisms
G06400QJ	Gal(b1-3)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc-ol	GM4 Type 1 LN	Linkage defined saccharide	2079.76	Amniota Bifurcata Bilateria Calloselasma rhodostoma Calloselasma Chordata Colubroidea Craniata Crotalinae Deuterostomia Dipnotetrapodomorpha Episquamata Eukaryota Eumetazoa Euteleostomi Gnathostomata Lepidosauria Metazoa Opisthokonta Sarcopterygii Sauria Sauropsida Serpentes Squamata Teleostomi Tetrapoda Toxicofera Unidentata Vertebrata Viperidae cellular organisms
G08005PG	Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-4)]Man(a1-3)[Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc-ol	GM4 Type 1 LN	Linkage defined saccharide	3683.31	Amniota Bifurcata Bilateria Calloselasma rhodostoma Calloselasma Chordata Colubroidea Craniata Crotalinae Deuterostomia Dipnotetrapodomorpha Episquamata Eukaryota Eumetazoa Euteleostomi Gnathostomata Lepidosauria Metazoa Opisthokonta Sarcopterygii Sauria Sauropsida Serpentes Squamata Teleostomi Tetrapoda Toxicofera Unidentata Vertebrata Viperidae cellular organisms
G08176ZW	Gal(a1-3)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-	Lewis x Lex N-acetyllactosamine SSEA-1 Type 2 LN a-Gal antigen	Linkage defined saccharide	691.25	Amniota Bifurcata Bilateria Chordata Colubroidea Craniata Deuterostomia Dipnotetrapodomorpha Elapidae Elapinae Episquamata Eukaryota Eumetazoa Euteleostomi Gnathostomata Lepidosauria Metazoa Naja kaouthia Naja Opisthokonta Sarcopterygii Sauria Sauropsida Serpentes Squamata Teleostomi Tetrapoda Toxicofera Unidentata Vertebrata cellular organisms
G11485OM	GalNAc(b1-4)GlcNAc(b1-2)Man(a1-3)[GalNAc(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]Man-ol	LDN LacdiNAc	Linkage defined saccharide	1829.69	Amniota Bifurcata Bilateria Bothrops moojeni Bothrops Chordata Colubroidea Craniata Crotalinae Deuterostomia Dipnotetrapodomorpha Episquamata Eukaryota Eumetazoa Euteleostomi Gnathostomata Lepidosauria Metazoa Opisthokonta Sarcopterygii Sauria Sauropsida Serpentes Squamata Teleostomi Tetrapoda Toxicofera Unidentata Vertebrata Viperidae cellular organisms
G11960UF	GalNAc(b1-4)GlcNAc(b1-2)Man(a1-3)[GalNAc(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	LDN LacdiNAc N-Glycan complex	Linkage defined saccharide	1722.65	Amniota Artiodactyla Bifurcata Bilateria Boreoeutheria Bos taurus Bos Bothrops moojeni Bothrops Bovidae Bovinae Catarrhini Chordata Colubroidea Craniata Crotalinae Deuterostomia Dipnotetrapodomorpha Episquamata Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Lepidosauria Mammalia Metazoa Opisthokonta Pecora Primates Ruminantia Sarcopterygii Sauria Sauropsida Serpentes Simiiformes Squamata Teleostomi Tetrapoda Theria Toxicofera Unidentata Vertebrata Viperidae cellular organisms
G14899VS	Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	GM4 N-Glycan complex Type 1 LN	Linkage defined saccharide	2368.84	Amniota Bifurcata Bilateria Calloselasma rhodostoma Calloselasma Chordata Colubroidea Craniata Crotalinae Deuterostomia Dipnotetrapodomorpha Episquamata Eukaryota Eumetazoa Euteleostomi Gnathostomata Lepidosauria Metazoa Opisthokonta Sarcopterygii Sauria Sauropsida Serpentes Squamata Teleostomi Tetrapoda Toxicofera Unidentata Vertebrata Viperidae cellular organisms
G14942ZU	Neu5Ac(a2-3)GalNAc(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	GM4 LDN LacdiNAc N-Glycan complex Type 1 LN	Linkage defined saccharide	2409.87	Amniota Bifurcata Bilateria Calloselasma rhodostoma Calloselasma Chordata Colubroidea Craniata Crotalinae Deuterostomia Dipnotetrapodomorpha Episquamata Eukaryota Eumetazoa Euteleostomi Gnathostomata Lepidosauria Metazoa Opisthokonta Sarcopterygii Sauria Sauropsida Serpentes Squamata Teleostomi Tetrapoda Toxicofera Unidentata Vertebrata Viperidae cellular organisms
G16147FU	Neu5Ac(a2-3)Gal(b1-3/4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-3/4)GlcNAc(b1-4)]Man(a1-3)[Neu5Ac(a2-3)Gal(b1-3/4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-3/4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc-ol	Polysialic acid Type 1 LN	Linkage defined saccharide	3683.31	Amniota Bifurcata Bilateria Calloselasma rhodostoma Calloselasma Chordata Colubroidea Craniata Crotalinae Deuterostomia Dipnotetrapodomorpha Episquamata Eukaryota Eumetazoa Euteleostomi Gnathostomata Lepidosauria Metazoa Opisthokonta Sarcopterygii Sauria Sauropsida Serpentes Squamata Teleostomi Tetrapoda Toxicofera Unidentata Vertebrata Viperidae cellular organisms
G20173ZW	GalNAc(b1-4)GlcNAc(b1-2)Man(a1-3)[Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	LDN LacdiNAc N-Glycan hybrid	Linkage defined saccharide	1640.59	Actinopteri Actinopterygii Amniota Bifurcata Bilateria Bothrops moojeni Bothrops Chordata Clupeocephala Colubroidea Craniata Crotalinae Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Dipnotetrapodomorpha Episquamata Eukaryota Eumetazoa Euteleostomi Gnathostomata Lepidosauria Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Sarcopterygii Sauria Sauropsida Serpentes Squamata Teleostei Teleostomi Tetrapoda Toxicofera Unidentata Vertebrata Viperidae cellular organisms
G21219OJ	Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-4)]Man(a1-3)[Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	GM4 N-Glycan complex Type 1 LN	Linkage defined saccharide	3025.07	Amniota Bifurcata Bilateria Calloselasma rhodostoma Calloselasma Chordata Colubroidea Craniata Crotalinae Deuterostomia Dipnotetrapodomorpha Episquamata Eukaryota Eumetazoa Euteleostomi Gnathostomata Lepidosauria Metazoa Opisthokonta Sarcopterygii Sauria Sauropsida Serpentes Squamata Teleostomi Tetrapoda Toxicofera Unidentata Vertebrata Viperidae cellular organisms
G21627CA	Neu5Ac(a2-3)GalNAc(b1-4)GlcNAc(b1-2)Man(a1-3)[GalNAc(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	LDN LacdiNAc N-Glycan complex	Linkage defined saccharide	2159.8	Amniota Bifurcata Bilateria Bothrops moojeni Bothrops Chordata Colubroidea Craniata Crotalinae Deuterostomia Dipnotetrapodomorpha Episquamata Eukaryota Eumetazoa Euteleostomi Gnathostomata Lepidosauria Metazoa Opisthokonta Sarcopterygii Sauria Sauropsida Serpentes Squamata Teleostomi Tetrapoda Toxicofera Unidentata Vertebrata Viperidae cellular organisms
G26179TW	Gal(b1-3)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	GM4 N-Glycan complex Type 1 LN	Linkage defined saccharide	2077.75	Amniota Bifurcata Bilateria Calloselasma rhodostoma Calloselasma Chordata Colubroidea Craniata Crotalinae Deuterostomia Dipnotetrapodomorpha Episquamata Eukaryota Eumetazoa Euteleostomi Gnathostomata Lepidosauria Metazoa Opisthokonta Sarcopterygii Sauria Sauropsida Serpentes Squamata Teleostomi Tetrapoda Toxicofera Unidentata Vertebrata Viperidae cellular organisms
G30447NS	Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	GM4 N-Glycan complex Type 1 LN	Linkage defined saccharide	3228.15	Amniota Bifurcata Bilateria Calloselasma rhodostoma Calloselasma Chordata Colubroidea Craniata Crotalinae Deuterostomia Dipnotetrapodomorpha Episquamata Eukaryota Eumetazoa Euteleostomi Gnathostomata Lepidosauria Metazoa Opisthokonta Sarcopterygii Sauria Sauropsida Serpentes Squamata Teleostomi Tetrapoda Toxicofera Unidentata Vertebrata Viperidae cellular organisms
G33178BD	Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc-ol	GM4 Type 1 LN	Linkage defined saccharide	3230.16	Amniota Bifurcata Bilateria Calloselasma rhodostoma Calloselasma Chordata Colubroidea Craniata Crotalinae Deuterostomia Dipnotetrapodomorpha Episquamata Eukaryota Eumetazoa Euteleostomi Gnathostomata Lepidosauria Metazoa Opisthokonta Sarcopterygii Sauria Sauropsida Serpentes Squamata Teleostomi Tetrapoda Toxicofera Unidentata Vertebrata Viperidae cellular organisms
G34383AG	GalNAc(b1-4)GlcNAc(b1-2)Man(a1-3)[Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	LDN LacdiNAc N-Glycan hybrid	Linkage defined saccharide	1786.65	Amniota Bifurcata Bilateria Bothrops moojeni Bothrops Chordata Colubroidea Craniata Crotalinae Deuterostomia Dipnotetrapodomorpha Episquamata Eukaryota Eumetazoa Euteleostomi Gnathostomata Lepidosauria Metazoa Opisthokonta Sarcopterygii Sauria Sauropsida Serpentes Squamata Teleostomi Tetrapoda Toxicofera Unidentata Vertebrata Viperidae cellular organisms
G34996GA	Man(a1-3)[Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-2/6)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc-ol	GM4 Type 1 LN	Linkage defined saccharide	1714.63	Amniota Bifurcata Bilateria Calloselasma rhodostoma Calloselasma Chordata Colubroidea Craniata Crotalinae Deuterostomia Dipnotetrapodomorpha Episquamata Eukaryota Eumetazoa Euteleostomi Gnathostomata Lepidosauria Metazoa Opisthokonta Sarcopterygii Sauria Sauropsida Serpentes Squamata Teleostomi Tetrapoda Toxicofera Unidentata Vertebrata Viperidae cellular organisms
G37845OW		GM4 N-Glycan complex Type 1 LN	Linkage defined saccharide	1915.69	Amniota Bifurcata Bilateria Calloselasma rhodostoma Calloselasma Chordata Colubroidea Craniata Crotalinae Deuterostomia Dipnotetrapodomorpha Episquamata Eukaryota Eumetazoa Euteleostomi Gnathostomata Lepidosauria Metazoa Opisthokonta Sarcopterygii Sauria Sauropsida Serpentes Squamata Teleostomi Tetrapoda Toxicofera Unidentata Vertebrata Viperidae cellular organisms
G38373BD	GalNAc(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	GM4 LDN LacdiNAc N-Glycan complex Type 1 LN	Linkage defined saccharide	2118.77	Amniota Bifurcata Bilateria Calloselasma rhodostoma Calloselasma Chordata Colubroidea Craniata Crotalinae Deuterostomia Dipnotetrapodomorpha Episquamata Eukaryota Eumetazoa Euteleostomi Gnathostomata Lepidosauria Metazoa Opisthokonta Sarcopterygii Sauria Sauropsida Serpentes Squamata Teleostomi Tetrapoda Toxicofera Unidentata Vertebrata Viperidae cellular organisms
G38625SV	Neu5Ac(a2-3)GalNAc(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	GM4 LDN LacdiNAc N-Glycan complex Type 1 LN	Linkage defined saccharide	2263.81	Amniota Bifurcata Bilateria Calloselasma rhodostoma Calloselasma Chordata Colubroidea Craniata Crotalinae Deuterostomia Dipnotetrapodomorpha Episquamata Eukaryota Eumetazoa Euteleostomi Gnathostomata Lepidosauria Metazoa Opisthokonta Sarcopterygii Sauria Sauropsida Serpentes Squamata Teleostomi Tetrapoda Toxicofera Unidentata Vertebrata Viperidae cellular organisms
G39023AU	Gal(b1-3)[Fuc(a1-4)]GlcNAc(b1-	Lea Lewis a Type 1 LN	Linkage defined saccharide	529.2	Amniota Artiodactyla Artverviricota Bacteria Bifurcata Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Campylobacterales Campylobacterota Canidae Caniformia Canis lupus familiaris Canis lupus Canis Carnivora Catarrhini Cercopithecidae Cercopithecinae Cercopithecoidea Chlorocebus aethiops Chlorocebus Chordata Colubroidea Craniata Deuterostomia Dipnotetrapodomorpha Elapidae Elapinae Episquamata Epsilonproteobacteria Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Felidae Feliformia Felinae Felis catus Felis Glires Gnathostomata Haplorrhini Helicobacter pylori Helicobacter Helicobacteraceae Hominidae Homininae Hominoidea Homo sapiens Homo Human immunodeficiency virus 1 Human immunodeficiency virus Lagomorpha Laurasiatheria Lentivirus Lepidosauria Leporidae Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Naja kaouthia Naja Opisthokonta Ortervirales Orthoretrovirinae Oryctolagus cuniculus Oryctolagus Pararnavirae Pecora Primate lentivirus group Primates Rattus Retroviridae Revtraviricetes Riboviria Rodentia Ruminantia Sarcopterygii Sauria Sauropsida Serpentes Simiiformes Squamata Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Toxicofera Unidentata Vertebrata Viruses cellular organisms unclassified Primate lentivirus group
G39453SO		N-acetyllactosamine Type 2 LN			Amniota Artiodactyla Bacteria Bifurcata Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Campylobacterales Campylobacterota Catarrhini Chordata Colubroidea Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dipnotetrapodomorpha Elapidae Elapinae Episquamata Epsilonproteobacteria Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Helicobacter pylori 26695 Helicobacter pylori Helicobacter Helicobacteraceae Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Lepidosauria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Naja kaouthia Naja Opisthokonta Pecora Primates Rodentia Ruminantia Sarcopterygii Sauria Sauropsida Serpentes Simiiformes Squamata Teleostomi Tetrapoda Theria Toxicofera Unidentata Vertebrata cellular organisms
G39852JB		N-acetyllactosamine Type 2 LN			Amniota Artiodactyla Bifurcata Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Colubroidea Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dipnotetrapodomorpha Elapidae Elapinae Episquamata Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Lepidosauria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Naja Opisthokonta Pecora Primates Rodentia Ruminantia Sarcopterygii Sauria Sauropsida Serpentes Simiiformes Squamata Teleostomi Tetrapoda Theria Toxicofera Unidentata Vertebrata cellular organisms
G41821MR	Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	Lewis x Lex N-Glycan complex N-acetyllactosamine SSEA-1 Type 2 LN	Linkage defined saccharide	2078.77	Amniota Archelosauria Archosauria Articulavirales Aves Bifurcata Bilateria Boreoeutheria Catarrhini Chordata Coelurosauria Colubroidea Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dinosauria Dipnotetrapodomorpha Elapidae Elapinae Episquamata Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Haploviricotina Hominidae Homininae Hominoidea Homo sapiens Homo Insthoviricetes Lepidosauria Mammalia Metazoa Monjiviricetes Mononegavirales Muroidea Myomorpha Naja kaouthia Naja Negarnaviricota Neognathae Opisthokonta Orthomyxoviridae Orthoparamyxovirinae Orthornavirae Paramyxoviridae Phasianidae Phasianinae Polyploviricotina Primates Respirovirus muris Respirovirus Riboviria Rodentia Sarcopterygii Sauria Saurischia Sauropsida Serpentes Simiiformes Squamata Teleostomi Tetrapoda Theria Theropoda Toxicofera Unidentata Vertebrata Viruses cellular organisms unclassified Orthomyxoviridae unidentified influenza virus
G42049JU	Neu5Ac(a2-3)Gal(b1-4)GlcNAc	GM4	Linkage defined saccharide	674.24	Amniota Artiodactyla Bacteria Bifurcata Bilateria Boreoeutheria Calloselasma rhodostoma Calloselasma Campylobacterales Campylobacterota Catarrhini Chordata Colubroidea Craniata Crotalinae Deuterostomia Dipnotetrapodomorpha Episquamata Epsilonproteobacteria Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Helicobacter pylori Helicobacter Helicobacteraceae Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Lepidosauria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Primates Rattus Rodentia Sarcopterygii Sauria Sauropsida Serpentes Simiiformes Squamata Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Toxicofera Unidentata Vertebrata Viperidae cellular organisms