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Glycoconjugate Repository

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Glycomics MS Repository for glycan annotations from GlycoWorkbench

LM-GlycoRepo

Repository for lectin-assisted multimodality data

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Guidelines

MIRAGE

mucin 4

Summary

Gene Symbol

Muc4

Organism

Mus musculus (house mouse)

External Links

NCBI Gene

140474

PubChem

140474

Alliance of Genome Resources

MGI:2153525

Annotation

Keyword

Cell adhesion
Cell membrane
Disulfide bond
EGF-like domain
Glycoprotein
Proteomics identification
Reference proteome
Repeat
Secreted
Signal
Transmembrane helix

Proteins

Displaying **all 2** entries
UniProt	Protein Name
A0A7N9VSW1
Q8JZM8	Ascites sialoglycoprotein Pancreatic adenocarcinoma mucin Testis mucin

Annotation information from UniProt.

Gene Ontology (GO)

Displaying entries **1 - 5** of 7 in total

1

2

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GO Term	Evidence Code	PMID
negative regulation of cell-matrix adhesion	ISO
hematopoietic progenitor cell differentiation	IMP	24029230
negative regulation of T cell mediated cytotoxicity directed against tumor cell target	ISO
cell-matrix adhesion	IEA
negative regulation of cell-cell adhesion	ISO

GO Hierarchy

biological process

cellular process [inference]

cell adhesion [inference]

cell-substrate adhesion [inference]

cell-matrix adhesion

cellular developmental process [inference]

cell differentiation [inference]

hematopoietic progenitor cell differentiation

multicellular organismal process [inference]

system process [inference]

digestive system process [inference]

maintenance of gastrointestinal epithelium

multicellular organismal-level homeostasis [inference]

anatomical structure homeostasis [inference]

tissue homeostasis [inference]

epithelial structure maintenance [inference]

maintenance of gastrointestinal epithelium

developmental process [inference]

cellular developmental process [inference]

cell differentiation [inference]

hematopoietic progenitor cell differentiation

homeostatic process [inference]

multicellular organismal-level homeostasis [inference]

anatomical structure homeostasis [inference]

tissue homeostasis [inference]

epithelial structure maintenance [inference]

maintenance of gastrointestinal epithelium

biological regulation [inference]

regulation of biological process [inference]

regulation of immune system process [inference]

negative regulation of immune system process [inference]

negative regulation of immune effector process [inference]

negative regulation of leukocyte mediated immunity [inference]

negative regulation of leukocyte mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of lymphocyte mediated immunity [inference]

negative regulation of T cell mediated immunity [inference]

negative regulation of T cell mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of T cell mediated immune response to tumor cell [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of immune response [inference]

negative regulation of adaptive immune response [inference]

negative regulation of adaptive immune response based on somatic recombination of immune receptors built from immunoglobulin superfamily domains [inference]

negative regulation of T cell mediated immunity [inference]

negative regulation of T cell mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of T cell mediated immune response to tumor cell [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of immune response to tumor cell [inference]

negative regulation of T cell mediated immune response to tumor cell [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

regulation of immune effector process [inference]

negative regulation of immune effector process [inference]

negative regulation of leukocyte mediated immunity [inference]

negative regulation of leukocyte mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of lymphocyte mediated immunity [inference]

negative regulation of T cell mediated immunity [inference]

negative regulation of T cell mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of T cell mediated immune response to tumor cell [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

regulation of leukocyte mediated immunity [inference]

regulation of leukocyte mediated cytotoxicity [inference]

negative regulation of leukocyte mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

regulation of T cell mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

regulation of T cell mediated cytotoxicity directed against tumor cell target [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of leukocyte mediated immunity [inference]

negative regulation of leukocyte mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of lymphocyte mediated immunity [inference]

negative regulation of T cell mediated immunity [inference]

negative regulation of T cell mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of T cell mediated immune response to tumor cell [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

regulation of lymphocyte mediated immunity [inference]

negative regulation of lymphocyte mediated immunity [inference]

negative regulation of T cell mediated immunity [inference]

negative regulation of T cell mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of T cell mediated immune response to tumor cell [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

regulation of T cell mediated immunity [inference]

regulation of T cell mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

regulation of T cell mediated cytotoxicity directed against tumor cell target [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of T cell mediated immunity [inference]

negative regulation of T cell mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of T cell mediated immune response to tumor cell [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

regulation of T cell mediated immune response to tumor cell [inference]

negative regulation of T cell mediated immune response to tumor cell [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

regulation of T cell mediated cytotoxicity directed against tumor cell target [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

regulation of immune response [inference]

regulation of adaptive immune response [inference]

negative regulation of adaptive immune response [inference]

negative regulation of adaptive immune response based on somatic recombination of immune receptors built from immunoglobulin superfamily domains [inference]

negative regulation of T cell mediated immunity [inference]

negative regulation of T cell mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of T cell mediated immune response to tumor cell [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

regulation of adaptive immune response based on somatic recombination of immune receptors built from immunoglobulin superfamily domains [inference]

regulation of T cell mediated immunity [inference]

regulation of T cell mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

regulation of T cell mediated cytotoxicity directed against tumor cell target [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of T cell mediated immunity [inference]

negative regulation of T cell mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of T cell mediated immune response to tumor cell [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

regulation of T cell mediated immune response to tumor cell [inference]

negative regulation of T cell mediated immune response to tumor cell [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

regulation of T cell mediated cytotoxicity directed against tumor cell target [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of adaptive immune response based on somatic recombination of immune receptors built from immunoglobulin superfamily domains [inference]

negative regulation of T cell mediated immunity [inference]

negative regulation of T cell mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of T cell mediated immune response to tumor cell [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

regulation of immune response to tumor cell [inference]

negative regulation of immune response to tumor cell [inference]

negative regulation of T cell mediated immune response to tumor cell [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

regulation of T cell mediated immune response to tumor cell [inference]

negative regulation of T cell mediated immune response to tumor cell [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

regulation of T cell mediated cytotoxicity directed against tumor cell target [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of immune response [inference]

negative regulation of adaptive immune response [inference]

negative regulation of adaptive immune response based on somatic recombination of immune receptors built from immunoglobulin superfamily domains [inference]

negative regulation of T cell mediated immunity [inference]

negative regulation of T cell mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of T cell mediated immune response to tumor cell [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of immune response to tumor cell [inference]

negative regulation of T cell mediated immune response to tumor cell [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of biological process [inference]

negative regulation of immune system process [inference]

negative regulation of immune effector process [inference]

negative regulation of leukocyte mediated immunity [inference]

negative regulation of leukocyte mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of lymphocyte mediated immunity [inference]

negative regulation of T cell mediated immunity [inference]

negative regulation of T cell mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of T cell mediated immune response to tumor cell [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of immune response [inference]

negative regulation of adaptive immune response [inference]

negative regulation of adaptive immune response based on somatic recombination of immune receptors built from immunoglobulin superfamily domains [inference]

negative regulation of T cell mediated immunity [inference]

negative regulation of T cell mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of T cell mediated immune response to tumor cell [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of immune response to tumor cell [inference]

negative regulation of T cell mediated immune response to tumor cell [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of cellular process [inference]

negative regulation of cell adhesion [inference]

negative regulation of cell-substrate adhesion [inference]

negative regulation of cell-matrix adhesion

negative regulation of cell-cell adhesion

negative regulation of cell killing [inference]

negative regulation of leukocyte mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of programmed cell death [inference]

negative regulation of apoptotic process

negative regulation of response to stimulus [inference]

negative regulation of response to biotic stimulus [inference]

negative regulation of response to tumor cell [inference]

negative regulation of immune response to tumor cell [inference]

negative regulation of T cell mediated immune response to tumor cell [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of immune response [inference]

negative regulation of adaptive immune response [inference]

negative regulation of adaptive immune response based on somatic recombination of immune receptors built from immunoglobulin superfamily domains [inference]

negative regulation of T cell mediated immunity [inference]

negative regulation of T cell mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of T cell mediated immune response to tumor cell [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of immune response to tumor cell [inference]

negative regulation of T cell mediated immune response to tumor cell [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

regulation of response to stimulus [inference]

regulation of response to biotic stimulus [inference]

negative regulation of response to biotic stimulus [inference]

negative regulation of response to tumor cell [inference]

negative regulation of immune response to tumor cell [inference]

negative regulation of T cell mediated immune response to tumor cell [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

regulation of response to tumor cell [inference]

negative regulation of response to tumor cell [inference]

negative regulation of immune response to tumor cell [inference]

negative regulation of T cell mediated immune response to tumor cell [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

regulation of immune response to tumor cell [inference]

negative regulation of immune response to tumor cell [inference]

negative regulation of T cell mediated immune response to tumor cell [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

regulation of T cell mediated immune response to tumor cell [inference]

negative regulation of T cell mediated immune response to tumor cell [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

regulation of T cell mediated cytotoxicity directed against tumor cell target [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of response to stimulus [inference]

negative regulation of response to biotic stimulus [inference]

negative regulation of response to tumor cell [inference]

negative regulation of immune response to tumor cell [inference]

negative regulation of T cell mediated immune response to tumor cell [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of immune response [inference]

negative regulation of adaptive immune response [inference]

negative regulation of adaptive immune response based on somatic recombination of immune receptors built from immunoglobulin superfamily domains [inference]

negative regulation of T cell mediated immunity [inference]

negative regulation of T cell mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of T cell mediated immune response to tumor cell [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of immune response to tumor cell [inference]

negative regulation of T cell mediated immune response to tumor cell [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

regulation of immune response [inference]

regulation of adaptive immune response [inference]

negative regulation of adaptive immune response [inference]

negative regulation of adaptive immune response based on somatic recombination of immune receptors built from immunoglobulin superfamily domains [inference]

negative regulation of T cell mediated immunity [inference]

negative regulation of T cell mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of T cell mediated immune response to tumor cell [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

regulation of adaptive immune response based on somatic recombination of immune receptors built from immunoglobulin superfamily domains [inference]

regulation of T cell mediated immunity [inference]

regulation of T cell mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

regulation of T cell mediated cytotoxicity directed against tumor cell target [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of T cell mediated immunity [inference]

negative regulation of T cell mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of T cell mediated immune response to tumor cell [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

regulation of T cell mediated immune response to tumor cell [inference]

negative regulation of T cell mediated immune response to tumor cell [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

regulation of T cell mediated cytotoxicity directed against tumor cell target [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of adaptive immune response based on somatic recombination of immune receptors built from immunoglobulin superfamily domains [inference]

negative regulation of T cell mediated immunity [inference]

negative regulation of T cell mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of T cell mediated immune response to tumor cell [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

regulation of immune response to tumor cell [inference]

negative regulation of immune response to tumor cell [inference]

negative regulation of T cell mediated immune response to tumor cell [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

regulation of T cell mediated immune response to tumor cell [inference]

negative regulation of T cell mediated immune response to tumor cell [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

regulation of T cell mediated cytotoxicity directed against tumor cell target [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of immune response [inference]

negative regulation of adaptive immune response [inference]

negative regulation of adaptive immune response based on somatic recombination of immune receptors built from immunoglobulin superfamily domains [inference]

negative regulation of T cell mediated immunity [inference]

negative regulation of T cell mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of T cell mediated immune response to tumor cell [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of immune response to tumor cell [inference]

negative regulation of T cell mediated immune response to tumor cell [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

regulation of cellular process [inference]

regulation of cell adhesion [inference]

negative regulation of cell adhesion [inference]

negative regulation of cell-substrate adhesion [inference]

negative regulation of cell-matrix adhesion

negative regulation of cell-cell adhesion

regulation of cell-substrate adhesion [inference]

regulation of cell-matrix adhesion [inference]

negative regulation of cell-matrix adhesion

negative regulation of cell-substrate adhesion [inference]

negative regulation of cell-matrix adhesion

regulation of cell-cell adhesion [inference]

negative regulation of cell-cell adhesion

regulation of cell killing [inference]

regulation of leukocyte mediated cytotoxicity [inference]

negative regulation of leukocyte mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

regulation of T cell mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

regulation of T cell mediated cytotoxicity directed against tumor cell target [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of cell killing [inference]

negative regulation of leukocyte mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

regulation of programmed cell death [inference]

regulation of apoptotic process [inference]

negative regulation of apoptotic process

negative regulation of programmed cell death [inference]

negative regulation of apoptotic process

negative regulation of cellular process [inference]

negative regulation of cell adhesion [inference]

negative regulation of cell-substrate adhesion [inference]

negative regulation of cell-matrix adhesion

negative regulation of cell-cell adhesion

negative regulation of cell killing [inference]

negative regulation of leukocyte mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity [inference]

negative regulation of T cell mediated cytotoxicity directed against tumor cell target

negative regulation of programmed cell death [inference]

negative regulation of apoptotic process

Displaying entries **1 - 5** of 8 in total

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GO Term	Evidence Code	PMID
extracellular region	IEA
extracellular space	ISO
extracellular space	IBA
plasma membrane	IEA
apical plasma membrane	ISO

GO Hierarchy

cellular component

protein-containing complex

cellular anatomical entity [inference]

membrane [inference]

plasma membrane

plasma membrane region [inference]

apical plasma membrane

cell projection membrane [inference]

microvillus membrane

external encapsulating structure [inference]

extracellular matrix [inference]

collagen-containing extracellular matrix

Displaying **all 3** entries
GO Term	Evidence Code	PMID
ErbB-2 class receptor binding	ISO
ErbB-2 class receptor binding	IBA
protein-containing complex binding	ISO

GO Hierarchy

molecular function

binding [inference]

protein binding [inference]

signaling receptor binding [inference]

ErbB-2 class receptor binding

protein-containing complex binding

Gene Ontology information from NCBI Gene and UniProt.

OrthoDB (Group)

Group level

Eukaryota

Group Name

notch

Functional Category

G: Carbohydrate transport and metabolism
M: Cell wall/membrane/envelope biogenesis
T: Signal transduction mechanisms

Displaying 1 entry
Gene Ontology
calcium ion binding

Displaying **all 3** entries
InterPro
EGF-like calcium-binding domain
EGF-like domain
EGF-type aspartate/asparagine hydroxylation site

Disease

Disease Ontology

Displaying 1 entry
DO ID	Disease Name	Source
DOID:0080599	Coronavirus infectious disease	Alliance of Genome Resources

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GlyCosmos Portal v4.5.0

Last updated: April 6, 2026