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Guidelines

MIRAGE

autophagy related 4B cysteine peptidase

Summary

Gene Symbol

ATG4B

Organism

Homo sapiens (human)

External Links

NCBI Gene

23192

PubChem

23192

Alliance of Genome Resources

HGNC:20790

JoGo

ATG4B

TogoVar

ATG4B

Annotation

Keyword

3D-structure
Acetylation
Alternative splicing
Autophagy
Cytoplasm
Cytoplasmic vesicle
Direct protein sequencing
Disulfide bond
Endoplasmic reticulum
Glycoprotein
Isopeptide bond
Mitochondrion
Phosphoprotein
Protein transport
Proteomics identification
Reference proteome
S-nitrosylation
Thiol protease
Ubl conjugation
Ubl conjugation pathway

Proteins

Displaying **all 3** entries
UniProt	Protein Name
B7ZA52
Q9Y4P1	AUT-like 1 cysteine endopeptidase Autophagy-related cysteine endopeptidase 1 Autophagy-related protein 4 homolog B
B3KVU2

Annotation information from UniProt.

Gene Ontology (GO)

Displaying entries **1 - 5** of 15 in total

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GO Term	Evidence Code	PMID
autophagosome assembly	IDA	21177865
autophagosome assembly	IBA
mitophagy	IMP	33773106
mitophagy	IBA
proteolysis	IDA	15169837 18387192 21177865

GO Hierarchy

biological process

metabolic process [inference]

nitrogen compound metabolic process [inference]

organonitrogen compound metabolic process [inference]

protein metabolic process [inference]

protein processing

protein modification process [inference]

protein delipidation

protein maturation [inference]

protein processing

catabolic process [inference]

cellular catabolic process [inference]

autophagy of mitochondrion [inference]

mitophagy

nucleophagy [inference]

piecemeal microautophagy of the nucleus

piecemeal microautophagy of the nucleus

cellular metabolic process [inference]

cellular catabolic process [inference]

autophagy of mitochondrion [inference]

mitophagy

nucleophagy [inference]

piecemeal microautophagy of the nucleus

piecemeal microautophagy of the nucleus

primary metabolic process [inference]

protein metabolic process [inference]

protein processing

protein modification process [inference]

protein delipidation

protein maturation [inference]

protein processing

organic substance metabolic process [inference]

macromolecule metabolic process [inference]

protein metabolic process [inference]

protein processing

protein modification process [inference]

protein delipidation

protein maturation [inference]

protein processing

macromolecule modification [inference]

protein modification process [inference]

protein delipidation

organonitrogen compound metabolic process [inference]

protein metabolic process [inference]

protein processing

protein modification process [inference]

protein delipidation

protein maturation [inference]

protein processing

cellular process [inference]

cellular metabolic process [inference]

cellular catabolic process [inference]

autophagy of mitochondrion [inference]

mitophagy

nucleophagy [inference]

piecemeal microautophagy of the nucleus

piecemeal microautophagy of the nucleus

cellular localization [inference]

cellular macromolecule localization [inference]

protein localization [inference]

protein localization to phagophore assembly site

establishment of protein localization [inference]

protein transport

process utilizing autophagic mechanism [inference]

autophagy of mitochondrion [inference]

mitophagy

nucleophagy [inference]

piecemeal microautophagy of the nucleus

piecemeal microautophagy of the nucleus

cellular component organization or biogenesis [inference]

cellular component organization [inference]

organelle organization [inference]

mitochondrion organization [inference]

autophagy of mitochondrion [inference]

mitophagy

vacuole organization [inference]

autophagosome organization [inference]

autophagosome assembly

organelle assembly [inference]

autophagosome assembly

organelle disassembly [inference]

autophagy of mitochondrion [inference]

mitophagy

cellular component disassembly [inference]

organelle disassembly [inference]

autophagy of mitochondrion [inference]

mitophagy

cellular component assembly [inference]

organelle assembly [inference]

autophagosome assembly

developmental process [inference]

anatomical structure development [inference]

tissue development [inference]

biomineral tissue development [inference]

otolith mineralization [inference]

otolith mineralization completed early in development

localization [inference]

macromolecule localization [inference]

cellular macromolecule localization [inference]

protein localization [inference]

protein localization to phagophore assembly site

establishment of protein localization [inference]

protein transport

establishment of localization [inference]

transport [inference]

organic substance transport [inference]

protein transport

nitrogen compound transport [inference]

protein transport

establishment of protein localization [inference]

protein transport

cellular localization [inference]

cellular macromolecule localization [inference]

protein localization [inference]

protein localization to phagophore assembly site

establishment of protein localization [inference]

protein transport

Displaying entries **1 - 5** of 6 in total

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GO Term	Evidence Code	PMID
autophagosome membrane	IDA	34562084
cytoplasm	IBA
mitochondrion	IEA
endoplasmic reticulum	IEA
cytosol	TAS

GO Hierarchy

cellular component

cellular anatomical entity [inference]

membrane [inference]

organelle membrane [inference]

bounding membrane of organelle [inference]

vacuolar membrane [inference]

autophagosome membrane

organelle [inference]

membrane-bounded organelle [inference]

vesicle [inference]

intracellular vesicle [inference]

cytoplasmic vesicle

intracellular membrane-bounded organelle [inference]

intracellular vesicle [inference]

cytoplasmic vesicle

intracellular organelle [inference]

intracellular membrane-bounded organelle [inference]

intracellular vesicle [inference]

cytoplasmic vesicle

Displaying entries **1 - 5** of 9 in total

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GO Term	Evidence Code	PMID
endopeptidase activity	IDA	18387192
cysteine-type endopeptidase activity	TAS
cysteine-type endopeptidase activity	IDA	21177865
cysteine-type endopeptidase activity	IBA
protein binding	IPI	15187094 16169070 18083104 19322194 20010802 20562859 21900206 25126726 30118682 32296183 33961781 34524948 35271311

GO Hierarchy

molecular function

catalytic activity [inference]

hydrolase activity [inference]

peptidase activity [inference]

endopeptidase activity

cysteine-type endopeptidase activity

cysteine-type peptidase activity

cysteine-type endopeptidase activity

protein-phosphatidylethanolamide deconjugating activity

catalytic activity, acting on a protein [inference]

peptidase activity [inference]

endopeptidase activity

cysteine-type endopeptidase activity

cysteine-type peptidase activity

cysteine-type endopeptidase activity

protein-phosphatidylethanolamide deconjugating activity

binding [inference]

protein binding

scaffold protein binding

Gene Ontology information from NCBI Gene and UniProt.

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GlyCosmos is a member of the GlySpace Alliance together with GlyGen and Glycomics@ExPASy.

Acknowledgements

Supported by JST NBDC Grant Number JPMJND2204

Partly supported by NIH Common Fund Grant #1U01GM125267-01

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GlyCosmos Portal v4.5.0

Last updated: April 6, 2026