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GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	August 18, 2024

Displaying entries **226 - 250** of **583** in total

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GlyTouCan ID ▲	IUPAC Condensed	Motifs	Subsumption Level	Monoisotopic Mass	Organisms
G39595FH			Glycosidic topology	3390.2	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Rattus norvegicus Rattus Riboviria Rodentia Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G39619TI			Glycosidic topology	2046.75	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Rattus norvegicus Rattus Riboviria Rodentia Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G39943KJ	GlcNAc(?1-?)Man(a1-3)[GlcNAc(?1-?)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan core basic	Linkage defined saccharide	1316.49	Acalyptratae Amniota Arthropoda Betacoronavirus Bilateria Boreoeutheria Brachycera Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Cyclorrhapha Deuterostomia Dicondylia Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Endopterygota Ephydroidea Eremoneura Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Mammalia Mandibulata Metazoa Muscomorpha Neoptera Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Panarthropoda Pancrustacea Pisoniviricetes Pisuviricota Primates Protostomia Pterygota Riboviria Sarbecovirus Sarcopterygii Schizophora Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Sophophora Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms melanogaster group melanogaster subgroup
G40198OD	Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan high mannose	Linkage defined saccharide	1558.54	50 kb inversion clade Acalypheae Acalyphoideae Acer pseudoplatanus Acer Acereae Aesculus hippocastanum Aesculus Amniota Amphiesmenoptera Amphipyrinae Anthemideae Archelosauria Archosauria Artemisia vulgaris Artemisia Artemisiinae Arthropoda Articulavirales Artiodactyla Artverviricota Ascomycota Aspergillaceae Aspergillus niger Aspergillus subgen. Circumdati Aspergillus Asteraceae Asterales Asteroideae Aves Betulaceae Bilateria Bombycidae Bombycinae Bombycoidea Bombyx mori Bombyx Boreoeutheria Bos taurus Bos Bovidae Bovinae Carnivora Carpinus betulus Carpinus Catarrhini Cavia porcellus Cavia Caviidae Chordata Coelurosauria Coturnix coturnix Coturnix Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dicondylia Dikarya Dinosauria Dipnotetrapodomorpha Discoba Ditrysia Ecdysozoa Embryophyta Endopterygota Euarchontoglires Euglenozoa Eukaryota Eumetazoa Euphorbiaceae Euphyllophyta Eurotiales Eurotiomycetes Eurotiomycetidae Euteleostomi Eutheria Fabaceae Fabales Fabeae Fagales Felidae Feliformia Felinae Felis catus Felis Friend murine leukemia virus Fungi Galliformes Galloanserae Gammaretrovirus Glires Glossata Gnathostomata Gunneridae Hadeninae Haplorrhini Heteroneura Hexapoda Hippocastaneae Hippocastanoideae Hologalegina Hominidae Homininae Hominoidea Homo sapiens Homo Hystricomorpha IRL clade Insecta Insthoviricetes Kinetoplastea Lamiales Laurasiatheria Lepidoptera Magnoliopsida Malpighiales Mamestra brassicae Mamestra Mammalia Mandibulata Mesangiospermae Metakinetoplastina Metazoa Muridae Murinae Murine leukemia virus Muroidea Mus musculus Mus Mus Myomorpha NPAAA clade Negarnaviricota Neognathae Neolepidoptera Neoptera Nicotiana alata Nicotiana Nicotianeae Nicotianoideae Noctuidae Noctuoidea Obtectomera Olea europaea Olea Oleaceae Oleeae Opisthokonta Ortervirales Orthomyxoviridae Orthoretrovirinae Orthornavirae Panarthropoda Pancrustacea Papilionoideae Pararnavirae Pecora Pentapetalae Perdicinae Pezizomycotina Phaseoleae Phaseolus vulgaris Phaseolus Phasianidae Pisum sativum Pisum Plusiinae Polyploviricotina Primates Protostomia Pterygota Rattus Retroviridae Revtraviricetes Riboviria Ricinus communis Ricinus Rodentia Ruminantia Sapindaceae Sapindales Sarcopterygii Sauria Saurischia Sauropsida Schizotrypanum Simiiformes Solanaceae Solanales Spermatophyta Spodoptera frugiperda Spodoptera Streptophyta Streptophytina Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Theropoda Tracheophyta Trichoplusia ni Trichoplusia Trypanosoma cruzi Trypanosoma Trypanosomatida Trypanosomatidae Vertebrata Viridiplantae Viruses asterids campanulids cellular organisms eudicotyledons fabids indigoferoid/millettioid clade lamiids leotiomyceta malvids rosids saccharomyceta unclassified Orthomyxoviridae unidentified influenza virus
G40206WX			Base composition with linkage	2493.94	Betacoronavirus Cornidovirineae Coronaviridae Nidovirales Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Riboviria Sarbecovirus Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Viruses
G40350YP	Glc(a1-2)Glc(a1-3)Glc(a1-3)Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(?1-		Linkage defined saccharide	1841.62	Amniota Articulavirales Bilateria Boreoeutheria Chordata Craniata Cricetidae Cricetinae Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Insthoviricetes Mammalia Mesocricetus auratus Mesocricetus Metazoa Muroidea Myomorpha Negarnaviricota Opisthokonta Orthomyxoviridae Orthornavirae Polyploviricotina Riboviria Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms unclassified Orthomyxoviridae unidentified influenza virus
G40792UP			Base composition with linkage	1811.68	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Rattus norvegicus Rattus Riboviria Rodentia Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G40926MX			Monosaccharide composition with linkage	2296.82	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Riboviria Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G41044JW			Monosaccharide composition with linkage	3390.2	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Rattus norvegicus Rattus Riboviria Rodentia Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G41126SR			Base composition with linkage	2443.9	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Rattus norvegicus Rattus Riboviria Rodentia Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G41247ZX			Base composition with linkage	1396.49	Amarillovirales Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Flasuviricetes Flaviviridae Glires Gnathostomata Haplorrhini Hepacivirus hominis Hepacivirus Hepatitis C virus (isolate H77) Hominidae Homininae Hominoidea Homo sapiens Homo Kitrinoviricota Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Rattus norvegicus Rattus Riboviria Rodentia Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms hepatitis C virus genotype 1a
G41821MR	Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	Lewis x Lex N-Glycan complex N-acetyllactosamine SSEA-1 Type 2 LN	Linkage defined saccharide	2078.77	Amniota Archelosauria Archosauria Articulavirales Aves Bifurcata Bilateria Boreoeutheria Catarrhini Chordata Coelurosauria Colubroidea Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dinosauria Dipnotetrapodomorpha Elapidae Elapinae Episquamata Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Haploviricotina Hominidae Homininae Hominoidea Homo sapiens Homo Insthoviricetes Lepidosauria Mammalia Metazoa Monjiviricetes Mononegavirales Muroidea Myomorpha Naja kaouthia Naja Negarnaviricota Neognathae Opisthokonta Orthomyxoviridae Orthoparamyxovirinae Orthornavirae Paramyxoviridae Phasianidae Phasianinae Polyploviricotina Primates Respirovirus muris Respirovirus Riboviria Rodentia Sarcopterygii Sauria Saurischia Sauropsida Serpentes Simiiformes Squamata Teleostomi Tetrapoda Theria Theropoda Toxicofera Unidentata Vertebrata Viruses cellular organisms unclassified Orthomyxoviridae unidentified influenza virus
G41882MT			Glycosidic topology	2661.95	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Riboviria Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G42028VS	Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	Lewis x Lex N-Glycan complex N-acetyllactosamine SSEA-1 Type 2 LN	Linkage defined saccharide	1786.65	Haploviricotina Monjiviricetes Mononegavirales Negarnaviricota Orthoparamyxovirinae Orthornavirae Paramyxoviridae Respirovirus muris Respirovirus Riboviria Viruses
G42358LZ	Neu5Ac(?2-?)Gal(?1-?)GlcNAc(?1-?)Man(a1-3)[Neu5Ac(?2-?)Gal(?1-?)GlcNAc(?1-?)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan core basic	Linkage defined saccharide	2222.78	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Riboviria Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G42864TB	Man(a1-3)[Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc-ol		Linkage defined saccharide	1195.42	Amniota Artiodactyla Avulavirinae Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haploviricotina Laurasiatheria Mammalia Metazoa Monjiviricetes Mononegavirales Negarnaviricota Opisthokonta Orthoavulavirus javaense Orthoavulavirus Orthornavirae Paramyxoviridae Riboviria Sarcopterygii Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata Viruses avian paramyxovirus 1 cellular organisms
G42962KI		N-Glycan core basic	Linkage defined saccharide	1932.71	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Riboviria Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G43157UW	Gal(?1-?)GlcNAc(?1-?)Man(a1-3)[GlcNAc(b1-4)][Gal(?1-?)GlcNAc(?1-?)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan core basic	Linkage defined saccharide	1843.67	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Riboviria Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G43417UB			Base composition with linkage	383.14	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Riboviria Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G43603TQ	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-4)]Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc-ol	Lewis x Lex N-acetyllactosamine SSEA-1 Type 2 LN	Linkage defined saccharide	2299.86	Articulavirales Insthoviricetes Negarnaviricota Orthomyxoviridae Orthornavirae Polyploviricotina Riboviria Viruses unclassified Orthomyxoviridae unidentified influenza virus
G43638QT			Monosaccharide composition with linkage	2483.9	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Riboviria Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G43642TU			Base composition with linkage	1754.66	Betacoronavirus Cornidovirineae Coronaviridae Nidovirales Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Riboviria Sarbecovirus Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Viruses
G43669FQ			Base composition with linkage	2370.86	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Riboviria Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G43734MM			Base composition with linkage	2411.88	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Riboviria Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G43769HG			Base composition with linkage	2046.75	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Riboviria Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms