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GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	August 18, 2024

Displaying entries **251 - 275** of **677** in total

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GlyTouCan ID	IUPAC Condensed	Motifs	Subsumption Level	Monoisotopic Mass	Organisms
G50757KG			Glycosidic topology	1868.7	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Rattus norvegicus Rattus Riboviria Rodentia Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G51287LK			Glycosidic topology	1608.6	Acalyptratae Amniota Arthropoda Betacoronavirus Bilateria Boreoeutheria Brachycera Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Cyclorrhapha Deuterostomia Dicondylia Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Endopterygota Ephydroidea Eremoneura Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Mammalia Mandibulata Metazoa Muscomorpha Neoptera Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Panarthropoda Pancrustacea Pisoniviricetes Pisuviricota Primates Protostomia Pterygota Riboviria Sarbecovirus Sarcopterygii Schizophora Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Sophophora Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms melanogaster group melanogaster subgroup
G51640FO			Glycosidic topology	2030.76	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Rattus norvegicus Rattus Riboviria Rodentia Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G51413EV			Glycosidic topology	2369.86	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Riboviria Rodentia Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G51795VM	Man(a1-3)[Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(?1-		Linkage defined saccharide	1031.35	Amniota Articulavirales Artiodactyla Artverviricota Ascomycota Aspergillaceae Aspergillus oryzae Aspergillus subgen. Circumdati Aspergillus Avulavirinae Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Carnivora Caryophyllaceae Caryophyllales Catarrhini Chordata Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dikarya Dipnotetrapodomorpha Embryophyta Euarchontoglires Eukaryota Eumetazoa Euphyllophyta Eurotiales Eurotiomycetes Eurotiomycetidae Euteleostomi Eutheria Felidae Feliformia Felinae Felis catus Felis Friend murine leukemia virus Fungi Gammaretrovirus Glires Gnathostomata Gunneridae Haplorrhini Haploviricotina Hominidae Homininae Hominoidea Homo sapiens Homo Human immunodeficiency virus Insthoviricetes Laurasiatheria Lentivirus Magnoliopsida Mammalia Mesangiospermae Metazoa Monjiviricetes Mononegavirales Muridae Murinae Murine leukemia virus Muroidea Mus musculus Mus Mus Myomorpha Negarnaviricota Opisthokonta Ortervirales Orthoavulavirus javaense Orthoavulavirus Orthomyxoviridae Orthoretrovirinae Orthornavirae Paramyxoviridae Pararnavirae Pecora Pentapetalae Pezizomycotina Polyploviricotina Primate lentivirus group Primates Rattus norvegicus Rattus Retroviridae Revtraviricetes Riboviria Rodentia Ruminantia Sarcopterygii Silene latifolia subsp. alba Silene latifolia Silene sect. Melandrium Silene subgen. Behenantha Silene Sileneae Simiiformes Solanaceae Solanales Solaneae Solanoideae Solanum lycopersicum Solanum subgen. Lycopersicon Solanum Spermatophyta Streptophyta Streptophytina Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Tracheophyta Vertebrata Viridiplantae Viruses asterids avian paramyxovirus 1 cellular organisms eudicotyledons lamiids leotiomyceta saccharomyceta unclassified Orthomyxoviridae unclassified Primate lentivirus group unclassified entries unclassified sequences unidentified influenza virus unidentified
G52527GH			Glycosidic topology	2587.92	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Riboviria Rodentia Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G52589SM			Glycosidic topology	2313.84	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Rattus norvegicus Rattus Riboviria Rodentia Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G53276NK	Neu5Ac(a2-?)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-?)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan complex Polysialic acid	Linkage defined saccharide	2222.78	Amniota Artverviricota Bilateria Blubervirales Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hepadnaviridae Hepatitis B virus Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Orthohepadnavirus Pararnavirae Primates Rattus norvegicus Rattus Revtraviricetes Riboviria Rodentia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G53631OG	Man(a1-2)Man(a1-3)[Man(a1-3)Glc(a1-6)]Glc(b1-4)GlcNAc(b1-4)GlcNAc(?1-		Linkage defined saccharide	1234.43	Articulavirales Insthoviricetes Negarnaviricota Orthomyxoviridae Orthornavirae Polyploviricotina Riboviria Viruses unclassified Orthomyxoviridae unidentified influenza virus
G54417MJ			Monosaccharide composition with linkage	2645.96	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Riboviria Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G54600FO			Monosaccharide composition with linkage	1915.69	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Rattus norvegicus Rattus Riboviria Rodentia Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G54612UD			Glycosidic topology	2263.81	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Riboviria Rodentia Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G54740VA			Glycosidic topology	3066.1	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Riboviria Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G55220VL	Man(a1-3)[Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan high mannose	Linkage defined saccharide	1234.43	Acalyptratae Amniota Archelosauria Archosauria Arthropoda Artiodactyla Aves Bacteria Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Brachycera Catarrhini Cavia porcellus Cavia Caviidae Chordata Coelurosauria Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Cyclorrhapha Deuterostomia Dicondylia Dinosauria Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Endopterygota Enterobacterales Enterobacteriaceae Ephydroidea Eremoneura Escherichia coli Escherichia Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Gammaproteobacteria Glires Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Hystricomorpha Insecta Laurasiatheria Mammalia Mandibulata Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha Neognathae Neoptera Opisthokonta Panarthropoda Pancrustacea Pecora Phasianidae Phasianinae Primates Protostomia Pseudomonadota Pterygota Rattus norvegicus Rattus Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Schizophora Simiiformes Sophophora Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Theropoda Vertebrata Viruses cellular organisms melanogaster group melanogaster subgroup
G55216FT			Base composition with linkage	2882.05	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Riboviria Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G55383ZG			Glycosidic topology	1567.58	Acalyptratae Amniota Arthropoda Betacoronavirus Bilateria Boreoeutheria Brachycera Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Cyclorrhapha Deuterostomia Dicondylia Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Endopterygota Ephydroidea Eremoneura Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Mammalia Mandibulata Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha Neoptera Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Panarthropoda Pancrustacea Pisoniviricetes Pisuviricota Primates Protostomia Pterygota Riboviria Rodentia Sarbecovirus Sarcopterygii Schizophora Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Sophophora Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms melanogaster group melanogaster subgroup
G55382TU			Base composition with linkage	1868.7	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Rattus norvegicus Rattus Riboviria Rodentia Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G55648EK	Man(a1-3)[Man(a1-2)Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc-ol		Linkage defined saccharide	1195.42	Avulavirinae Haploviricotina Monjiviricetes Mononegavirales Negarnaviricota Orthoavulavirus javaense Orthoavulavirus Orthornavirae Paramyxoviridae Riboviria Viruses avian paramyxovirus 1
G55868RH		N-Glycan core basic	Linkage defined saccharide	1681.62	Betacoronavirus Cornidovirineae Coronaviridae Nidovirales Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Riboviria Sarbecovirus Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Viruses
G56039JB	Neu5Ac(a2-3/6)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-4)][Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan hybrid Polysialic acid	Linkage defined saccharide	2093.74	Artverviricota Friend murine leukemia virus Gammaretrovirus Murine leukemia virus Ortervirales Orthoretrovirinae Pararnavirae Retroviridae Revtraviricetes Riboviria Viruses
G56299XO	Neu5Ac(a2-3)Gal(b1-3)GalNAc(?1-	GM4 O-Glycan core 1 fuzzy	Linkage defined saccharide	674.24	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dipnotetrapodomorpha Embecovirus Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Lagomorpha Leporidae Mammalia Metazoa Muridae Murinae Murine coronavirus Murine hepatitis virus strain A59 Murine hepatitis virus Muroidea Mus musculus Mus Mus Myomorpha Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Oryctolagus cuniculus Oryctolagus Pisoniviricetes Pisuviricota Primates Riboviria Rodentia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G56284ZY			Glycosidic topology	2160.82	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Riboviria Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G56610MH			Base composition with linkage	2030.76	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Riboviria Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G56791SN	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-4)][Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2354.86	Amniota Artiodactyla Artverviricota Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Equidae Equus caballus Equus Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Filoviridae Glires Gnathostomata Haplorrhini Haploviricotina Hominidae Homininae Hominoidea Homo sapiens Homo Human immunodeficiency virus Laurasiatheria Lentivirus Mammalia Metazoa Monjiviricetes Mononegavirales Muridae Murinae Muroidea Myomorpha Negarnaviricota Opisthokonta Ortervirales Orthomarburgvirus marburgense Orthomarburgvirus Orthoretrovirinae Orthornavirae Pararnavirae Pecora Perissodactyla Primate lentivirus group Primates Rattus norvegicus Rattus Retroviridae Revtraviricetes Riboviria Rodentia Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms unclassified Primate lentivirus group
G56903ZB	GlcNAc(?1-?)Man(a1-3)[GlcNAc(?1-?)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	N-Glycan core basic	Linkage defined saccharide	1462.54	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Riboviria Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms