GlyCosmos Portal

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GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	August 18, 2024

Displaying entries **351 - 375** of **391** in total

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GlyTouCan ID	IUPAC Condensed	Motifs	Subsumption Level	Monoisotopic Mass ▼	Organisms
G90719NY	Gal(b1-4)Gal(b1-3)GalNAc(a1-	O-Glycan core 1 T antigen TF antigen	Linkage defined saccharide	545.2	Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Deuterostomia Eukaryota Eumetazoa Euteleosteomorpha Euteleostomi Gnathostomata Metazoa Neopterygii Oncorhynchus mykiss Oncorhynchus Opisthokonta Osteoglossocephalai Protacanthopterygii Salmonidae Salmoniformes Salmoninae Teleostei Teleostomi Vertebrata cellular organisms
G97874DV	Neu5Gc(a2-6)GalNAc-ol		Linkage defined saccharide	530.2	Actinopteri Actinopterygii Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Clupeocephala Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleosteomorpha Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Neopterygii Oncorhynchus masou ishikawae Oncorhynchus masou Oncorhynchus mykiss Oncorhynchus Opisthokonta Osteoglossocephalai Pecora Primates Protacanthopterygii Ruminantia Salmonidae Salmoniformes Salmoninae Salvelinus leucomaenis japonicus Salvelinus leucomaenis pluvius Salvelinus leucomaenis Salvelinus Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G75206XT	Neu5Gc(a2-6)GalNAc(?1-		Linkage defined saccharide	528.18	Actinopteri Actinopterygii Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Clupeocephala Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleosteomorpha Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Neopterygii Opisthokonta Osteoglossocephalai Pecora Protacanthopterygii Ruminantia Salmonidae Salmoniformes Salmoninae Salvelinus leucomaenis pluvius Salvelinus leucomaenis Salvelinus Sarcopterygii Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G08371CD	Neu5Ac(a2-6)GalNAc(?1-		Linkage defined saccharide	512.19	Actinopteri Actinopterygii Amniota Amphibia Anura Artiodactyla Batrachia Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Cercopithecidae Cercopithecinae Cercopithecoidea Chordata Clupeocephala Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleosteomorpha Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Lithobates palustris Lithobates Macaca radiata Macaca Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neobatrachia Neopterygii Oncorhynchus mykiss Oncorhynchus Opisthokonta Osteoglossocephalai Pecora Primates Protacanthopterygii Ranidae Ranoidea Rattus norvegicus Rattus Rodentia Ruminantia Salmonidae Salmoniformes Salmoninae Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Vertebrata artificial sequences cellular organisms other entries other sequences synthetic construct
G53517XC	Neu5Gc(a2-3)Gal(?1-		Linkage defined saccharide	487.15	Actinopteri Actinopterygii Amniota Bilateria Boreoeutheria Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Dipnotetrapodomorpha Equidae Equus caballus Equus Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Perissodactyla Sarcopterygii Teleostei Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G04991IT	Kdn(a2-6)GalNAc-ol		Linkage defined saccharide	473.17	Actinopteri Actinopterygii Bilateria Botia Botiidae Chordata Clupeocephala Cobitoidei Craniata Cypriniformes Cypriniphysae Deuterostomia Eukaryota Eumetazoa Euteleostomi Gnathostomata Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Teleostei Teleostomi Vertebrata cellular organisms
G77824XV	Neu5Ac(a2-3)Gal(?1-		Linkage defined saccharide	471.16	Actinopteri Actinopterygii Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Pecora Primates Rodentia Ruminantia Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G30207PZ	Neu5Ac(a2-3)Gal(b1-	GM4	Linkage defined saccharide	471.16	Acanthomorphata Actinopteri Actinopterygii African cichlids Amniota Archelosauria Archosauria Artiodactyla Aves Balaenoptera acutorostrata Balaenoptera brydei Balaenoptera edeni Balaenoptera Balaenopteridae Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Cetacea Channichthyidae Chionodraco myersi Chionodraco Chordata Cichlidae Cichliformes Cichlomorphae Clupeocephala Coelurosauria Craniata Cricetidae Cricetinae Ctenosquamata Cyprinidae Cypriniformes Cyprininae Cypriniphysae Cyprinoidei Cyprinus carpio Cyprinus Delphinidae Delphinus delphis Delphinus Deuterostomia Dinosauria Dipnotetrapodomorpha Euacanthomorphacea Euarchontoglires Eukaryota Eumetazoa Eupercaria Eurypterygia Euteleosteomorpha Euteleostomi Eutheria Gadariae Gadidae Gadiformes Gadoidei Gadus morhua Gadus Galliformes Galloanserae Gallus gallus Gallus Glires Globicephala macrorhynchus Globicephala Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Leuciscidae Leuciscinae Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Mysticeti Neognathae Neopterygii Neoteleostei Nototheniidae Notothenioidei Odontoceti Opisthokonta Orcinus orca Orcinus Oreochromini Oreochromis mossambicus Oreochromis Ostariophysi Osteoglossocephalai Otomorpha Otophysi Ovalentaria Paracanthopterygii Pecora Perciformes Percomorphaceae Phasianidae Phasianinae Phocoenidae Phocoenoides dalli Phocoenoides Phodopus sungorus Phodopus Physeter catodon Physeter Physeteridae Primates Pseudocrenilabrinae Rattus Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Scardinius erythrophthalmus Scardinius Simiiformes Stenella coeruleoalba Stenella Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Theropoda Trematomus lepidorhinus Trematomus Tursiops truncatus Tursiops Vertebrata Whippomorpha Zeiogadaria cellular organisms
G78703NH	Kdn(a2-6)GalNAc(a1-	Tn antigen	Linkage defined saccharide	471.16	Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Cobitidae Cobitinae Cobitoidei Craniata Cypriniformes Cypriniphysae Deuterostomia Eukaryota Eumetazoa Euteleostomi Gnathostomata Metazoa Misgurnus anguillicaudatus Misgurnus Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Teleostei Teleostomi Vertebrata cellular organisms
G82625KX	Kdn(a2-3)Gal(?1-		Linkage defined saccharide	430.13	Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Eukaryota Eumetazoa Euteleostomi Gnathostomata Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Teleostei Teleostomi Vertebrata cellular organisms
G00031MO	Gal(b1-3)GalNAc(a1-	O-Glycan core 1 T antigen TF antigen	Linkage defined saccharide	383.14	Acalyptratae Acanthomorphata Actinopteri Actinopterygii Amniota Arthropoda Artiodactyla Bacteria Bacteroidaceae Bacteroidales Bacteroides ovatus Bacteroides Bacteroidia Bacteroidota/Chlorobiota group Bacteroidota Betacoronavirus Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Brachycera Canidae Caniformia Canis lupus familiaris Canis lupus Canis Carnivora Catarrhini Cercopithecidae Cercopithecinae Cercopithecoidea Chordata Clupeocephala Cornidovirineae Coronaviridae Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Ctenosquamata Cyclorrhapha Deuterostomia Dicondylia Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Endopterygota Enterobacterales Enterobacteriaceae Ephydroidea Eremoneura Euarchontoglires Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Eutheria FCB group Gadariae Gadidae Gadiformes Gadoidei Gadus chalcogrammus Gadus Gammaproteobacteria Glires Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Lagomorpha Laurasiatheria Leporidae Macaca mulatta Macaca Mammalia Mandibulata Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha Neoptera Neopterygii Neoteleostei Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Oryctolagus cuniculus Oryctolagus Osteoglossocephalai Panarthropoda Pancrustacea Paracanthopterygii Pecora Pisoniviricetes Pisuviricota Pollachius pollachius Pollachius Primates Protostomia Pseudomonadota Pterygota Rattus norvegicus Rattus Riboviria Rodentia Ruminantia Sarbecovirus Sarcopterygii Schizophora Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Shigella dysenteriae Shigella Simiiformes Sophophora Suidae Suina Sus scrofa domesticus Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Vertebrata Viruses Zeiogadaria cellular organisms melanogaster group melanogaster subgroup
G21469TE	Gal(b1-3)GlcNAc(a1-		Linkage defined saccharide	383.14	Acanthomorphata Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Ctenosquamata Deuterostomia Dissostichus mawsoni Dissostichus Euacanthomorphacea Eukaryota Eumetazoa Eupercaria Eurypterygia Euteleosteomorpha Euteleostomi Gnathostomata Metazoa Neopterygii Neoteleostei Nototheniidae Notothenioidei Opisthokonta Osteoglossocephalai Perciformes Percomorphaceae Teleostei Teleostomi Vertebrata cellular organisms
G74621DY	Gal(b1-4)Glc(?1-		Linkage defined saccharide	342.12	Actinopteri Actinopterygii Amniota Bilateria Boreoeutheria Catarrhini Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Primates Sarcopterygii Simiiformes Teleostei Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G40748TG			Linkage defined saccharide	254.1	Acanthomorphata Actinopteri Actinopterygii Amniota Bacteria Bilateria Boreoeutheria Catarrhini Chordata Clupeocephala Craniata Ctenosquamata Cyanobacteriota/Melainabacteria group Cyanobacteriota Cyanophyceae Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Eutheria Gadariae Gadidae Gadiformes Gadoidei Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Neopterygii Neoteleostei Opisthokonta Oscillatoriaceae Oscillatoriales Oscillatoriophycideae Osteoglossocephalai Paracanthopterygii Phormidium tenue Phormidium Pollachius pollachius Pollachius Primates Rattus Rodentia Sarcopterygii Simiiformes Teleostei Teleostomi Terrabacteria group Tetrapoda Theria Vertebrata Zeiogadaria cellular organisms
G38776RF	Hex(?1-				Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Eukaryota Eumetazoa Euteleostomi Gnathostomata Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Teleostei Teleostomi Vertebrata cellular organisms
G02379LL					Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Deuterostomia Eukaryota Eumetazoa Euteleosteomorpha Euteleostomi Gnathostomata Metazoa Neopterygii Oncorhynchus mykiss Oncorhynchus Opisthokonta Osteoglossocephalai Protacanthopterygii Salmonidae Salmoniformes Salmoninae Teleostei Teleostomi Vertebrata cellular organisms
G06367TC					Actinopteri Actinopterygii Amniota Archelosauria Archosauria Artiodactyla Aves Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Carassius auratus Carassius Catarrhini Cercopithecidae Cercopithecinae Cercopithecoidea Chlorocebus aethiops Chlorocebus Chordata Clupeocephala Coelurosauria Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Cyprinidae Cypriniformes Cyprininae Cypriniphysae Cyprinoidei Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neognathae Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Pecora Phasianidae Phasianinae Primates Rattus Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms
G08355EN					Actinopteri Actinopterygii Amniota Bacteria Betaproteobacteria Bilateria Boreoeutheria Chordata Clupeocephala Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleosteomorpha Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Neisseria meningitidis Neisseria Neisseriaceae Neisseriales Neopterygii Oncorhynchus mykiss Oncorhynchus Opisthokonta Osteoglossocephalai Protacanthopterygii Pseudomonadota Rattus Rodentia Salmonidae Salmoniformes Salmoninae Sarcopterygii Teleostei Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G10293PS		O-Glycan core 1 T antigen TF antigen			Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Deuterostomia Eukaryota Eumetazoa Euteleosteomorpha Euteleostomi Gnathostomata Metazoa Neopterygii Oncorhynchus mykiss Oncorhynchus Opisthokonta Osteoglossocephalai Protacanthopterygii Salmonidae Salmoniformes Salmoninae Teleostei Teleostomi Vertebrata cellular organisms
G19059PI					Acalyptratae Actinopteri Actinopterygii Amniota Ampullariidae Ampullarioidea Archelosauria Architaenioglossa Archosauria Arthropoda Artiodactyla Aves Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Brachycera Caenogastropoda Canidae Caniformia Canis lupus familiaris Canis lupus Canis Caprinae Carassius auratus Carassius Carnivora Catarrhini Cavia porcellus Cavia Caviidae Cercopithecidae Cercopithecinae Cercopithecoidea Cetacea Chondrichthyes Chordata Clupeocephala Coelurosauria Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Cyclorrhapha Cyprinidae Cypriniformes Cyprininae Cypriniphysae Cyprinoidei Deuterostomia Dicondylia Dinosauria Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Elasmobranchii Endopterygota Ephydroidea Eremoneura Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Felidae Feliformia Felinae Felis catus Felis Galliformes Galloanserae Gallus gallus Gallus Gastropoda Gerbillinae Glires Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Hystricomorpha Insecta Lagomorpha Laurasiatheria Leporidae Lophotrochozoa Macaca mulatta Macaca Mammalia Mandibulata Meriones unguiculatus Meriones Metazoa Mollusca Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha Neognathae Neoptera Neopterygii Opisthokonta Oryctolagus cuniculus Oryctolagus Ostariophysi Osteoglossocephalai Otomorpha Otophysi Ovis aries Ovis Panarthropoda Pancrustacea Pecora Phasianidae Phasianinae Pomacea Primates Protostomia Pterygota Rattus norvegicus Rattus Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Schizophora Simiiformes Sophophora Spiralia Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Theropoda Vertebrata Whippomorpha cellular organisms melanogaster group melanogaster subgroup
G32724PG					Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Deuterostomia Eukaryota Eumetazoa Euteleosteomorpha Euteleostomi Gnathostomata Metazoa Neopterygii Oncorhynchus mykiss Oncorhynchus Opisthokonta Osteoglossocephalai Protacanthopterygii Salmonidae Salmoniformes Salmoninae Teleostei Teleostomi Vertebrata cellular organisms
G15021LG	Glc(?1-				50 kb inversion clade Acanthomorphata Actinopteri Actinopterygii Amniota Archaea Bilateria Boreoeutheria Catarrhini Chordata Clupeocephala Craniata Ctenosquamata Deuterostomia Dipnotetrapodomorpha Embryophyta Euarchontoglires Eukaryota Eumetazoa Euphyllophyta Euryarchaeota Eurypterygia Euteleosteomorpha Euteleostomi Eutheria Fabaceae Fabales Gadariae Gadidae Gadiformes Gadoidei Glires Gnathostomata Gunneridae Halobacteria Haloferacaceae Haloferacales Haloferax volcanii Haloferax Haplorrhini Hologalegina Hominidae Homininae Hominoidea Homo sapiens Homo IRL clade Magnoliopsida Mammalia Medicago sativa Medicago Mesangiospermae Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha NPAAA clade Neopterygii Neoteleostei Opisthokonta Osteoglossocephalai Papilionoideae Paracanthopterygii Pentapetalae Pollachius pollachius Pollachius Primates Rodentia Sarcopterygii Simiiformes Spermatophyta Stenosarchaea group Streptophyta Streptophytina Teleostei Teleostomi Tetrapoda Theria Tracheophyta Trifolieae Vertebrata Viridiplantae Zeiogadaria cellular organisms eudicotyledons fabids rosids
G68158BT	Gal(?1-				Acanthomorphata Actinopteri Actinopterygii Amniota Bilateria Boreoeutheria Catarrhini Chordata Clupeocephala Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Ctenosquamata Deuterostomia Dipnotetrapodomorpha Embryophyta Euarchontoglires Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Eutheria Gadariae Gadidae Gadiformes Gadoidei Gadus chalcogrammus Gadus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muroidea Myomorpha Neopterygii Neoteleostei Opisthokonta Osteoglossocephalai Paracanthopterygii Pollachius pollachius Pollachius Primates Rodentia Sarcopterygii Simiiformes Streptophyta Streptophytina Teleostei Teleostomi Tetrapoda Theria Vertebrata Viridiplantae Zeiogadaria cellular organisms
G89166NZ					Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Deuterostomia Eukaryota Eumetazoa Euteleosteomorpha Euteleostomi Gnathostomata Metazoa Neopterygii Oncorhynchus mykiss Oncorhynchus Opisthokonta Osteoglossocephalai Protacanthopterygii Salmonidae Salmoniformes Salmoninae Teleostei Teleostomi Vertebrata cellular organisms
G23110MM					Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Deuterostomia Eukaryota Eumetazoa Euteleosteomorpha Euteleostomi Gnathostomata Metazoa Neopterygii Oncorhynchus masou ishikawae Oncorhynchus masou Oncorhynchus Opisthokonta Osteoglossocephalai Protacanthopterygii Salmonidae Salmoniformes Salmoninae Salvelinus leucomaenis pluvius Salvelinus leucomaenis Salvelinus namaycush Salvelinus Teleostei Teleostomi Vertebrata cellular organisms