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GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	August 18, 2024

Displaying entries **351 - 375** of **458** in total

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GlyTouCan ID	IUPAC Condensed	Motifs	Subsumption Level	Monoisotopic Mass ▲	Organisms
G60230HH	Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan high mannose	Linkage defined saccharide	1882.64	50 kb inversion clade Acalyptratae Amniota Arachnida Arthropoda Ascomycota Bilateria Boreoeutheria Brachycera Catarrhini Chelicerata Chordata Craniata Cyclorrhapha Deuterostomia Dicondylia Dikarya Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Embryophyta Endopterygota Ephydroidea Eremoneura Euarchontoglires Eukaryota Eumetazoa Euphyllophyta Euteleostomi Eutheria Fabaceae Fabales Fungi Glires Glycine max Glycine subgen. Soja Glycine Gnathostomata Gunneridae Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Magnoliopsida Mammalia Mandibulata Mesangiospermae Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha NPAAA clade Neoptera Opisthokonta Panarthropoda Pancrustacea Papilionoideae Pentapetalae Phaseoleae Phaseolus vulgaris Phaseolus Primates Protostomia Pterygota Rattus norvegicus Rattus Rodentia Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales Saccharomycetes Saccharomycotina Sarcopterygii Schizophora Scorpiones Simiiformes Sophophora Spermatophyta Streptophyta Streptophytina Teleostomi Tetrapoda Theria Tracheophyta Vertebrata Viridiplantae cellular organisms eudicotyledons fabids indigoferoid/millettioid clade melanogaster group melanogaster subgroup rosids saccharomyceta
G70101JE			Glycosidic topology	1882.64	Acalyptratae Amniota Arthropoda Ascomycota Betacoronavirus Bilateria Boreoeutheria Brachycera Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Cyclorrhapha Deuterostomia Dicondylia Dikarya Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Endopterygota Ephydroidea Eremoneura Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Fungi Glires Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Mammalia Mandibulata Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha Neoptera Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Panarthropoda Pancrustacea Pisoniviricetes Pisuviricota Primates Protostomia Pterygota Rattus norvegicus Rattus Riboviria Rodentia Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales Saccharomycetes Saccharomycotina Sarbecovirus Sarcopterygii Schizophora Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Sophophora Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms melanogaster group melanogaster subgroup saccharomyceta
G66257KQ	Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc-ol		Linkage defined saccharide	1884.66	50 kb inversion clade Acalyptratae Aculeata Alphainfluenzavirus influenzae Alphainfluenzavirus Amniota Anthophila Apidae Apinae Apini Apis mellifera Apis Apocrita Apoidea Arthropoda Articulavirales Artiodactyla Artverviricota Bilateria Boreoeutheria Bovidae Brachycera Caprinae Catarrhini Chordata Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Cyclorrhapha Deuterostomia Dicondylia Dipnotetrapodomorpha Diptera Discoba Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Embryophyta Endopterygota Ephydroidea Eremoneura Euarchontoglires Euglenozoa Eukaryota Eumetazoa Euphyllophyta Euteleostomi Eutheria Fabaceae Fabales Fabeae Glires Glycine max Glycine subgen. Soja Glycine Gnathostomata Gunneridae Haplorrhini Hexapoda Hologalegina Hominidae Homininae Hominoidea Homo sapiens Homo Human immunodeficiency virus Hymenoptera IRL clade Influenza A virus Insecta Insthoviricetes Kinetoplastea Laurasiatheria Lens culinaris Lens Lentivirus Magnoliopsida Mammalia Mandibulata Mesangiospermae Metakinetoplastina Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha NPAAA clade Negarnaviricota Neoptera Opisthokonta Ortervirales Orthomyxoviridae Orthoretrovirinae Orthornavirae Ovis aries Ovis Panarthropoda Pancrustacea Papilionoideae Pararnavirae Pecora Pentapetalae Phaseoleae Phaseolus coccineus Phaseolus lunatus Phaseolus vulgaris Phaseolus Polyploviricotina Primate lentivirus group Primates Protostomia Pterygota Rattus Retroviridae Revtraviricetes Riboviria Rodentia Ruminantia Sarcopterygii Schizophora Simiiformes Sophophora Spermatophyta Streptophyta Streptophytina Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Tracheophyta Trypanosoma brucei Trypanosoma Trypanosomatida Trypanosomatidae Trypanozoon Vertebrata Viridiplantae Viruses cellular organisms eudicotyledons fabids indigoferoid/millettioid clade melanogaster group melanogaster subgroup rosids unclassified Orthomyxoviridae unclassified Primate lentivirus group unidentified influenza virus
G03842KQ			Glycosidic topology	1907.68	Acalyptratae Amniota Arthropoda Bilateria Boreoeutheria Brachycera Catarrhini Chordata Craniata Cyclorrhapha Deuterostomia Dicondylia Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Endopterygota Ephydroidea Eremoneura Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Mammalia Mandibulata Metazoa Muscomorpha Neoptera Opisthokonta Panarthropoda Pancrustacea Primates Protostomia Pterygota Sarcopterygii Schizophora Simiiformes Sophophora Teleostomi Tetrapoda Theria Vertebrata cellular organisms melanogaster group melanogaster subgroup
G35731IU	Man(a1-3)[Gal4Me(b1-3)[Gal3Me(b1-6)]GalNAc(b1-4)GlcNAc(b1-2)Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	LDN LacdiNAc N-Glycan core basic	Linkage defined saccharide	1946.72	Bilateria Eukaryota Eumetazoa Eupulmonata Euthyneura Gastropoda Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Lophotrochozoa Metazoa Mollusca Opisthokonta Panpulmonata Protostomia Spiralia Stylommatophora cellular organisms
G07016SY	Man(a1-3)[Gal4Me(b1-3)[Gal3Me(b1-6)]GalNAc(b1-4)GlcNAc(b1-2)Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc-ol	LDN LacdiNAc	Linkage defined saccharide	1948.74	Bilateria Eukaryota Eumetazoa Eupulmonata Euthyneura Gastropoda Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Lophotrochozoa Metazoa Mollusca Opisthokonta Panpulmonata Protostomia Spiralia Stylommatophora cellular organisms
G26038CK	Gal3Me(b1-6)Gal3Me(b1-3)[Gal3Me(b1-6)]GlcNAc(b1-4)GlcNAc(b1-2)Man(a1-3)[Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan core basic	Linkage defined saccharide	1976.73	Bilateria Eukaryota Eumetazoa Eupulmonata Euthyneura Gastropoda Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Lophotrochozoa Metazoa Mollusca Opisthokonta Panpulmonata Protostomia Spiralia Stylommatophora cellular organisms
G32687NU	Gal3Me(b1-6)Gal3Me(b1-3)[Gal3Me(b1-6)]GalNAc(b1-4)GlcNAc(b1-2)Man(a1-3)[Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	LDN LacdiNAc N-Glycan core basic	Linkage defined saccharide	1976.73	Bilateria Eukaryota Eumetazoa Eupulmonata Euthyneura Gastropoda Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Lophotrochozoa Metazoa Mollusca Opisthokonta Panpulmonata Protostomia Spiralia Stylommatophora cellular organisms
G82142CI	Gal3Me(b1-6)Gal3Me(b1-3)[Gal3Me(b1-6)]GalNAc(b1-4)GlcNAc(b1-2)Man(a1-3)[Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)GlcNAc	LDN LacdiNAc N-Glycan core basic	Linkage defined saccharide	1976.73	Bilateria Eukaryota Eumetazoa Eupulmonata Euthyneura Gastropoda Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Lophotrochozoa Metazoa Mollusca Opisthokonta Panpulmonata Protostomia Spiralia Stylommatophora cellular organisms
G67951MU	Gal3Me(b1-6)Gal3Me(b1-3)[Gal3Me(b1-6)]GalNAc(b1-4)GlcNAc(b1-2)Man(a1-3)[Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)GlcNAc-ol	LDN LacdiNAc	Linkage defined saccharide	1978.75	Bilateria Eukaryota Eumetazoa Eupulmonata Euthyneura Gastropoda Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Lophotrochozoa Metazoa Mollusca Opisthokonta Panpulmonata Protostomia Spiralia Stylommatophora cellular organisms
G43897GC	Glc(a1-3)Glc(a1-3)Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc		Linkage defined saccharide	2003.67	Aedes albopictus Aedes Aedini Arthropoda Bilateria Culicidae Culicinae Culicoidea Culicomorpha Dicondylia Diptera Ecdysozoa Endopterygota Eukaryota Eumetazoa Hexapoda Insecta Mandibulata Metazoa Nematocera Neoptera Opisthokonta Panarthropoda Pancrustacea Protostomia Pterygota Stegomyia cellular organisms
G82018TQ	Glc(a1-3)Glc(a1-3)Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(?1-		Linkage defined saccharide	2003.67	Aedes albopictus Aedes Aedini Amniota Arthropoda Bilateria Boreoeutheria Chordata Craniata Cricetidae Cricetinae Culicidae Culicinae Culicoidea Culicomorpha Deuterostomia Dicondylia Dipnotetrapodomorpha Diptera Ecdysozoa Endopterygota Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Hexapoda Insecta Mammalia Mandibulata Mesocricetus auratus Mesocricetus Metazoa Muroidea Myomorpha Nematocera Neoptera Opisthokonta Panarthropoda Pancrustacea Protostomia Pterygota Rodentia Sarcopterygii Stegomyia Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G28449FY	Gal3Me(b1-6)Gal3Me(b1-3)[Gal3Me(b1-6)Gal3Me(b1-6)]GlcNAc(b1-4)GlcNAc(b1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan core basic	Linkage defined saccharide	2020.76	Bilateria Eukaryota Eumetazoa Eupulmonata Euthyneura Gastropoda Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Lophotrochozoa Metazoa Mollusca Opisthokonta Panpulmonata Protostomia Spiralia Stylommatophora cellular organisms
G42304BW	Gal3Me(b1-6)Gal3Me(b1-3)[Gal3Me(b1-6)Gal3Me(b1-6)]GalNAc(b1-4)GlcNAc(b1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	LDN LacdiNAc N-Glycan core basic	Linkage defined saccharide	2020.76	Bilateria Eukaryota Eumetazoa Eupulmonata Euthyneura Gastropoda Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Lophotrochozoa Metazoa Mollusca Opisthokonta Panpulmonata Protostomia Spiralia Stylommatophora cellular organisms
G72668YK	Gal3Me(b1-6)Gal3Me(b1-3)[Gal3Me(b1-6)Gal3Me(b1-6)]GalNAc(b1-4)GlcNAc(b1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	LDN LacdiNAc N-Glycan core basic	Linkage defined saccharide	2020.76	Bilateria Eukaryota Eumetazoa Eupulmonata Euthyneura Gastropoda Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Lophotrochozoa Metazoa Mollusca Opisthokonta Panpulmonata Protostomia Spiralia Stylommatophora cellular organisms
G82207VX	Glc(a1-3)Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan high mannose	Linkage defined saccharide	2044.7	50 kb inversion clade Acrididae Acrididea Acridoidea Acridomorpha Alpharhabdovirinae Amniota Amphiesmenoptera Amylomyces rouxii Amylomyces Arthropoda Artiodactyla Ascomycota Aspergillaceae Aspergillus niger Aspergillus subgen. Circumdati Aspergillus Asterias amurensis Asterias rubens Asterias Asteriidae Asteroidea Asterozoa BOP clade Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Caelifera Canavalia ensiformis Canavalia Catarrhini Chordata Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dicondylia Dikarya Dipnotetrapodomorpha Ditrysia Ecdysozoa Echinodermata Eleutherozoa Embryophyta Endopterygota Euarchontoglires Eukaryota Eumetazoa Euphyllophyta Eurotiales Eurotiomycetes Eurotiomycetidae Euteleostomi Eutheria Fabaceae Fabales Forcipulatacea Forcipulatida Fungi incertae sedis Fungi Glires Glossata Gnathostomata Gunneridae Haplorrhini Haploviricotina Heteroneura Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Laurasiatheria Lepidoptera Liliopsida Locusta migratoria Locusta Magnoliopsida Mammalia Mandibulata Mesangiospermae Metazoa Monjiviricetes Mononegavirales Mucoraceae Mucorales Mucorineae Mucoromycetes Mucoromycota Mucoromycotina Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha NPAAA clade Negarnaviricota Neolepidoptera Neoptera Noctuidae Noctuoidea Obtectomera Oedipodinae Opisthokonta Orthoptera Orthornavirae Oryza sativa Oryza Oryzeae Oryzinae Oryzoideae Panarthropoda Pancrustacea Papilionoideae Pecora Pentapetalae Petrosaviidae Pezizomycotina Phaseoleae Phaseolus vulgaris Phaseolus Plusiinae Poaceae Poales Polyneoptera Primates Protostomia Pterygota Rhabdoviridae Riboviria Rodentia Ruminantia Sarcopterygii Simiiformes Spermatophyta Streptophyta Streptophytina Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Tracheophyta Trichoplusia ni Trichoplusia Vertebrata Vesicular stomatitis virus Vesiculovirus Viridiplantae Viruses cellular organisms commelinids eudicotyledons fabids indigoferoid/millettioid clade leotiomyceta rosids saccharomyceta unclassified Vesiculovirus unclassified entries unclassified sequences unidentified
G12464FX		N-Glycan core basic	Linkage defined saccharide	2044.7	Acalyptratae Arthropoda Bilateria Brachycera Cyclorrhapha Dicondylia Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Endopterygota Ephydroidea Eremoneura Eukaryota Eumetazoa Hexapoda Insecta Mandibulata Metazoa Muscomorpha Neoptera Opisthokonta Panarthropoda Pancrustacea Protostomia Pterygota Schizophora Sophophora cellular organisms melanogaster group melanogaster subgroup
G10997HT	Gal3Me(b1-6)Gal3Me(b1-3)[Gal3Me(b1-6)]GalNAc(b1-4)GlcNAc(b1-2)Man(a1-3)[Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	LDN LacdiNAc N-Glycan core basic	Linkage defined saccharide	2122.79	Bilateria Eukaryota Eumetazoa Eupulmonata Euthyneura Gastropoda Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Lophotrochozoa Metazoa Mollusca Opisthokonta Panpulmonata Protostomia Spiralia Stylommatophora cellular organisms
G37009XT	Man(a1-3)[Gal3Me(b1-6)Gal3Me(b1-3)[Gal3Me(b1-6)]GlcNAc(b1-4)GlcNAc(b1-2)Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan core basic	Linkage defined saccharide	2122.79	Bilateria Eukaryota Eumetazoa Eupulmonata Euthyneura Gastropoda Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Lophotrochozoa Metazoa Mollusca Opisthokonta Panpulmonata Protostomia Spiralia Stylommatophora cellular organisms
G43499RV	Man(a1-3)[Gal3Me(b1-6)Gal3Me(b1-3)[Gal3Me(b1-6)]GalNAc(b1-4)GlcNAc(b1-2)Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	LDN LacdiNAc N-Glycan core basic	Linkage defined saccharide	2122.79	Bilateria Eukaryota Eumetazoa Eupulmonata Euthyneura Gastropoda Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Lophotrochozoa Metazoa Mollusca Opisthokonta Panpulmonata Protostomia Spiralia Stylommatophora cellular organisms
G64310FP	Gal3Me(b1-6)Gal3Me(b1-3)[Gal3Me(b1-6)]GalNAc(b1-4)GlcNAc(b1-2)Man(a1-3)[Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	LDN LacdiNAc N-Glycan core basic	Linkage defined saccharide	2122.79	Bilateria Eukaryota Eumetazoa Eupulmonata Euthyneura Gastropoda Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Lophotrochozoa Metazoa Mollusca Opisthokonta Panpulmonata Protostomia Spiralia Stylommatophora cellular organisms
G82902UU	Man(a1-3)[Gal3Me(b1-6)Gal3Me(b1-3)[Gal3Me(b1-6)]GalNAc(b1-4)GlcNAc(b1-2)Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	LDN LacdiNAc N-Glycan core basic	Linkage defined saccharide	2122.79	Bilateria Eukaryota Eumetazoa Eupulmonata Euthyneura Gastropoda Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Lophotrochozoa Metazoa Mollusca Opisthokonta Panpulmonata Protostomia Spiralia Stylommatophora cellular organisms
G79779NX	Gal3Me(b1-6)Gal3Me(b1-3)[Gal3Me(b1-6)]GlcNAc(b1-4)GlcNAc(b1-2)Man(a1-3)[Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan core basic	Linkage defined saccharide	2122.79	Bilateria Eukaryota Eumetazoa Eupulmonata Euthyneura Gastropoda Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Lophotrochozoa Metazoa Mollusca Opisthokonta Panpulmonata Protostomia Spiralia Stylommatophora cellular organisms
G44088KZ	Man(a1-3)[Gal3Me(b1-6)Gal3Me(b1-3)[Gal3Me(b1-6)]GalNAc(b1-4)GlcNAc(b1-2)Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc-ol	LDN LacdiNAc	Linkage defined saccharide	2124.81	Bilateria Eukaryota Eumetazoa Eupulmonata Euthyneura Gastropoda Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Lophotrochozoa Metazoa Mollusca Opisthokonta Panpulmonata Protostomia Spiralia Stylommatophora cellular organisms
G92249IB	Gal3Me(b1-6)Gal3Me(b1-3)[Gal3Me(b1-6)]GalNAc(b1-4)GlcNAc(b1-2)Man(a1-3)[Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc-ol	LDN LacdiNAc	Linkage defined saccharide	2124.81	Bilateria Eukaryota Eumetazoa Eupulmonata Euthyneura Gastropoda Helicidae Helicina Helicoidea Helix pomatia Helix Heterobranchia Lophotrochozoa Metazoa Mollusca Opisthokonta Panpulmonata Protostomia Spiralia Stylommatophora cellular organisms