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GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	August 18, 2024

Displaying entries **3951 - 3975** of **8356** in total

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GlyTouCan ID	IUPAC Condensed	Motifs	Subsumption Level	Monoisotopic Mass ▲	Organisms
G80583XN	Fuc(a1-2)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	Blood group H (type 2) Leb Lewis b Ley N-Glycan core basic	Linkage defined saccharide	1713.63	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G03930BU			Base composition with linkage	1713.63	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Primates Rattus norvegicus Rattus Rodentia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G50915WK	Man(a1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc-ol	Polysialic acid	Linkage defined saccharide	1714.63	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G58809AD	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc-ol	N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1714.63	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G79732PC	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3/6)[Man(a1-3/6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc-ol	Polysialic acid	Linkage defined saccharide	1714.63	Amniota Artiodactyla Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Sarcopterygii Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G97183VL			Glycosidic topology	1718.47	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G35380EO	GalNAc?S(b1-4)GlcNAc(b1-2)Man(a1-3)[Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan hybrid	Linkage defined saccharide	1720.55	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G00645SW	GalNAc3/4S(b1-4)GlcNAc(b1-2)Man(a1-3)[Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan hybrid	Linkage defined saccharide	1720.55	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G13363OF	GalNAc3/4S(b1-4)GlcNAc(b1-2)Man(a1-3)[Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan hybrid	Linkage defined saccharide	1720.55	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G45681QN	GalNAc4S(b1-4)GlcNAc(b1-2)Man(a1-3)[Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	4O-sulfated LDN N-Glycan hybrid	Linkage defined saccharide	1720.55	Amniota Artiodactyla Bilateria Boreoeutheria Bovidae Caprinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Ovis aries Ovis Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G86385RC	Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan high mannose	Linkage defined saccharide	1720.59	50 kb inversion clade Abreae Abrus precatorius Abrus Acalypheae Acalyphoideae Acalyptratae Acrogymnospermae Aesculus hippocastanum Aesculus Alpharhabdovirinae Amniota Amphiesmenoptera Amphipyrinae Anserineae Anthemideae Archelosauria Archosauria Artemisia vulgaris Artemisia Artemisiinae Arthropoda Articulavirales Artiodactyla Ascomycota Aspergillaceae Aspergillus niger Aspergillus subgen. Circumdati Aspergillus Asteraceae Asterales Asteroideae Aves BOP clade Batoidea Betula pubescens Betula Betulaceae Bilateria Bombycidae Bombycinae Bombycoidea Bombyx mori Bombyx Boreoeutheria Bos taurus Bos Bovidae Bovinae Brachycera Carpinus betulus Carpinus Caryophyllales Catarrhini Cavia porcellus Cavia Caviidae Chenopodiaceae Chenopodioideae Chondrichthyes Chordata Coelurosauria Conifers I Coturnix coturnix Coturnix Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Cyclorrhapha Deuterostomia Dicondylia Dikarya Dinosauria Dipnotetrapodomorpha Diptera Discoba Ditrysia Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Elasmobranchii Embryophyta Endopterygota Ephydroidea Eremoneura Euarchontoglires Euglenozoa Eukaryota Eumetazoa Euphorbiaceae Euphyllophyta Eurotiales Eurotiomycetes Eurotiomycetidae Euteleostomi Eutheria Fabaceae Fabales Fabeae Fagales Fungi Galliformes Galloanserae Glires Glossata Gnathostomata Gunneridae Hadeninae Haplorrhini Haploviricotina Heteroneura Hexapoda Hippocastaneae Hippocastanoideae Hologalegina Hominidae Homininae Hominoidea Homo sapiens Homo Hystricomorpha IRL clade Insecta Insthoviricetes Kinetoplastea Komagataella pastoris Komagataella Lamiales Laurasiatheria Lepidoptera Liliopsida Magnoliopsida Malpighiales Mamestra brassicae Mamestra Mammalia Mandibulata Mesangiospermae Metakinetoplastina Metazoa Monjiviricetes Mononegavirales Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha NPAAA clade Negarnaviricota Neognathae Neolepidoptera Neoptera Noctuidae Noctuoidea Obtectomera Olea europaea Olea Oleaceae Oleeae Opisthokonta Orthomyxoviridae Orthornavirae Oryza sativa Oryza Oryzeae Oryzinae Oryzoideae Panarthropoda Pancrustacea Papilionoideae Pecora Pentapetalae Perdicinae Petrosaviidae Pezizomycotina Phaffomycetaceae Phaseoleae Phaseolus vulgaris Phaseolus Phasianidae Pinaceae Pinales Pinidae Pinopsida Pinus heldreichii Pinus subgen. Pinus Pinus Pisum sativum Pisum Plusiinae Poaceae Poales Polyploviricotina Primates Protostomia Pterygota Rattus Rhabdoviridae Riboviria Ricinus communis Ricinus Rodentia Ruminantia Saccharomycetales Saccharomycetes Saccharomycotina Sapindaceae Sapindales Sarcopterygii Sauria Saurischia Sauropsida Schizophora Schizotrypanum Simiiformes Sophophora Spermatophyta Spinacia oleracea Spinacia Spodoptera frugiperda Spodoptera Streptophyta Streptophytina Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Tetronarce californica Tetronarce Theria Theropoda Torpedinidae Torpediniformes Tracheophyta Trichoplusia ni Trichoplusia Trypanosoma brucei Trypanosoma cruzi Trypanosoma Trypanosomatida Trypanosomatidae Trypanozoon Vertebrata Vesicular stomatitis virus Vesiculovirus Viridiplantae Viruses asterids campanulids cellular organisms commelinids eudicotyledons fabids indigoferoid/millettioid clade lamiids leotiomyceta malvids melanogaster group melanogaster subgroup rosids saccharomyceta unclassified Orthomyxoviridae unclassified Vesiculovirus unidentified influenza virus
G10601XM		N-Glycan core basic	Linkage defined saccharide	1720.59	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G21359LU	Glc(a1-3)Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-6)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan core basic	Linkage defined saccharide	1720.59	Amniota Artiodactyla Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Sarcopterygii Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G31429YW	Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan high mannose	Linkage defined saccharide	1720.59	Acalypheae Acalyphoideae Amniota Amphiesmenoptera Arthropoda Artiodactyla Bilateria Bombycidae Bombycinae Bombycoidea Bombyx mori Bombyx Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dicondylia Dipnotetrapodomorpha Ditrysia Ecdysozoa Embryophyta Endopterygota Euarchontoglires Eukaryota Eumetazoa Euphorbiaceae Euphyllophyta Euteleostomi Eutheria Glires Glossata Gnathostomata Gunneridae Hadeninae Haplorrhini Haploviricotina Heteroneura Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Laurasiatheria Lepidoptera Magnoliopsida Malpighiales Mamestra brassicae Mamestra Mammalia Mandibulata Mesangiospermae Metazoa Monjiviricetes Mononegavirales Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Negarnaviricota Neolepidoptera Neoptera Noctuidae Noctuoidea Obtectomera Opisthokonta Orthoparamyxovirinae Orthornavirae Panarthropoda Pancrustacea Paramyxoviridae Pecora Pentapetalae Primates Protostomia Pterygota Respirovirus muris Respirovirus Riboviria Ricinus communis Ricinus Rodentia Ruminantia Sarcopterygii Simiiformes Spermatophyta Streptophyta Streptophytina Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Tracheophyta Vertebrata Viridiplantae Viruses cellular organisms eudicotyledons fabids rosids
G34314SG	Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-3)[Man(a1-2)Man(a1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan core basic	Linkage defined saccharide	1720.59	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G35732LD	Glc(a1-3)Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan high mannose	Linkage defined saccharide	1720.59	Amniota Archelosauria Archosauria Artiodactyla Aves Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Laurasiatheria Mammalia Metazoa Neognathae Opisthokonta Pecora Phasianidae Phasianinae Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms
G08060JF	Glc(a1-3)Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-6)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan core basic	Linkage defined saccharide	1720.59	Amniota Artiodactyla Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Sarcopterygii Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G14548ZL		N-Glycan high mannose	Linkage defined saccharide	1720.59	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Rattus norvegicus Rattus Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G22477EB	Man(a1-3)[Man(a1-2)Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan high mannose	Linkage defined saccharide	1720.59	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Rattus Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G26335RK		N-Glycan core basic	Linkage defined saccharide	1720.59	Acalyptratae Amniota Arthropoda Bilateria Boreoeutheria Brachycera Catarrhini Chordata Craniata Cyclorrhapha Deuterostomia Dicondylia Dipnotetrapodomorpha Diptera Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Endopterygota Ephydroidea Eremoneura Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Mammalia Mandibulata Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha Neoptera Opisthokonta Panarthropoda Pancrustacea Primates Protostomia Pterygota Rodentia Sarcopterygii Schizophora Simiiformes Sophophora Teleostomi Tetrapoda Theria Vertebrata cellular organisms melanogaster group melanogaster subgroup
G32149KV	Glc(a1-3)Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan high mannose	Linkage defined saccharide	1720.59	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G34567KI	Man(a1-2)Man(a1-?)[Man(a1-2)Man(a1-?)]Man(a1-?)[Man(a1-2)Man(a1-?)]Man(a1-4)GlcNAc(b1-4)GlcNAc(b1-		Linkage defined saccharide	1720.59	Amniota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G39111QK	Man(a1-3)[Man(a1-2)Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan high mannose	Linkage defined saccharide	1720.59	Amniota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Rattus norvegicus Rattus Rodentia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G40702WU	Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan high mannose	Linkage defined saccharide	1720.59	50 kb inversion clade Amniota Archelosauria Archosauria Artiodactyla Ascomycota Aves Batoidea Bilateria Boreoeutheria Catarrhini Cavia porcellus Cavia Caviidae Chondrichthyes Chordata Coelurosauria Craniata Deuterostomia Dikarya Dinosauria Dipnotetrapodomorpha Elasmobranchii Embryophyta Euarchontoglires Eukaryota Eumetazoa Euphyllophyta Euteleostomi Eutheria Fabaceae Fabales Fungi Galliformes Galloanserae Glires Gnathostomata Gunneridae Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Hystricomorpha Laurasiatheria Magnoliopsida Mammalia Mesangiospermae Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha NPAAA clade Neognathae Opisthokonta Papilionoideae Pentapetalae Phaseoleae Phaseolus vulgaris Phaseolus Primates Rodentia Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales Saccharomycetes Saccharomycotina Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Spermatophyta Streptophyta Streptophytina Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Tetronarce californica Tetronarce Theria Theropoda Torpedinidae Torpediniformes Tracheophyta Vertebrata Viridiplantae cellular organisms eudicotyledons fabids indigoferoid/millettioid clade rosids saccharomyceta
G41824PC	Glc(a1-2)Glc(a1-3)Glc(a1-3)Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan core basic	Linkage defined saccharide	1720.59	Alveolata Amniota Ascomycota Bilateria Boreoeutheria Chordata Ciliophora Craniata Deuterostomia Dikarya Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Fungi Glires Gnathostomata Hymenostomatida Intramacronucleata Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Oligohymenophorea Opisthokonta Rodentia Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales Saccharomycetes Saccharomycotina Sar Sarcopterygii Teleostomi Tetrahymena pyriformis Tetrahymena Tetrahymenidae Tetrahymenina Tetrapoda Theria Vertebrata cellular organisms saccharomyceta