GlyCosmos Portal

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GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	August 18, 2024

Displaying entries **376 - 400** of **465** in total

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GlyTouCan ID	IUPAC Condensed	Motifs	Subsumption Level ▲	Monoisotopic Mass	Organisms
G93839ET	Neu5Ac(a2-3)GalNAc(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	GM4 LDN LacdiNAc N-Glycan complex Type 1 LN	Linkage defined saccharide	3066.1	Amniota Bifurcata Bilateria Calloselasma rhodostoma Calloselasma Chordata Colubroidea Craniata Crotalinae Deuterostomia Dipnotetrapodomorpha Episquamata Eukaryota Eumetazoa Euteleostomi Gnathostomata Lepidosauria Metazoa Opisthokonta Sarcopterygii Sauria Sauropsida Serpentes Squamata Teleostomi Tetrapoda Toxicofera Unidentata Vertebrata Viperidae cellular organisms
G93694YU	Neu5Ac(a2-8)Neu5Ac(a2-3)[Fuc(a1-2)Gal(b1-3)GalNAc(b1-4)]Gal(b1-4)Glc(b1-	2-Fucosyl-GD1b	Linkage defined saccharide	1435.5	Amniota Archelosauria Archosauria Artiodactyla Aves Bilateria Boreoeutheria Catarrhini Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gliridae Glirinae Glis glis Glis Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Neognathae Opisthokonta Phasianidae Phasianinae Primates Rattus Rodentia Sarcopterygii Sauria Saurischia Sauropsida Sciuromorpha Simiiformes Suidae Suina Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms
G94027HD	GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-4)][GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1884.7	Amniota Archelosauria Archosauria Aves Bilateria Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Metazoa Neognathae Opisthokonta Phasianidae Phasianinae Sarcopterygii Sauria Saurischia Sauropsida Teleostomi Tetrapoda Theropoda Vertebrata cellular organisms
G94106MV	Man(a1-3)[Man(a1-6)]Man(a1-6)Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-		Linkage defined saccharide	1072.38	Amniota Amphibia Anura Archelosauria Archosauria Aves Batrachia Bilateria Boreoeutheria Catarrhini Chordata Coelurosauria Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Lithobates pipiens Lithobates Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neobatrachia Neognathae Opisthokonta Phasianidae Phasianinae Primates Rana Ranidae Ranoidea Rodentia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms
G95223JW	Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(?1-		Linkage defined saccharide	1031.35	Amniota Archelosauria Archosauria Artiodactyla Aves Bilateria Boreoeutheria Bovidae Caprinae Carnivora Catarrhini Chordata Coelurosauria Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Felidae Feliformia Felinae Felis catus Felis Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muroidea Myomorpha Neognathae Opisthokonta Ovis aries Ovis Pecora Phasianidae Phasianinae Primates Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms unclassified entries unclassified sequences unidentified
G95440AP	Neu5Ac(a2-3)GalNAc(b1-4)GlcNAc(b1-	LDN LacdiNAc	Linkage defined saccharide	715.26	Amniota Bifurcata Bilateria Calloselasma rhodostoma Calloselasma Chordata Colubroidea Craniata Crotalinae Deuterostomia Dipnotetrapodomorpha Episquamata Eukaryota Eumetazoa Euteleostomi Gnathostomata Lepidosauria Metazoa Opisthokonta Sarcopterygii Sauria Sauropsida Serpentes Squamata Teleostomi Tetrapoda Toxicofera Unidentata Vertebrata Viperidae cellular organisms
G95462VZ	Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc-ol	GM4 Type 1 LN	Linkage defined saccharide	2573.94	Amniota Bifurcata Bilateria Calloselasma rhodostoma Calloselasma Chordata Colubroidea Craniata Crotalinae Deuterostomia Dipnotetrapodomorpha Episquamata Eukaryota Eumetazoa Euteleostomi Gnathostomata Lepidosauria Metazoa Opisthokonta Sarcopterygii Sauria Sauropsida Serpentes Squamata Teleostomi Tetrapoda Toxicofera Unidentata Vertebrata Viperidae cellular organisms
G95782RB	GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-4)][GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex	Linkage defined saccharide	1925.72	Amniota Archelosauria Archosauria Aves Bilateria Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Metazoa Neognathae Opisthokonta Phasianidae Phasianinae Sarcopterygii Sauria Saurischia Sauropsida Teleostomi Tetrapoda Theropoda Vertebrata cellular organisms unclassified entries unclassified sequences unidentified
G95910WJ	Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	GM4 N-Glycan complex Type 1 LN	Linkage defined saccharide	2571.92	Amniota Bifurcata Bilateria Calloselasma rhodostoma Calloselasma Chordata Colubroidea Craniata Crotalinae Deuterostomia Dipnotetrapodomorpha Episquamata Eukaryota Eumetazoa Euteleostomi Gnathostomata Lepidosauria Metazoa Opisthokonta Sarcopterygii Sauria Sauropsida Serpentes Squamata Teleostomi Tetrapoda Toxicofera Unidentata Vertebrata Viperidae cellular organisms
G96709CX	Neu5Ac(a2-8)Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-3)Gal(b1-4)Glc(b1-	GM4 Lactotriaosylceramide Lc3 Type 1 LN	Linkage defined saccharide	1289.44	Amniota Archelosauria Archosauria Aves Bilateria Boreoeutheria Catarrhini Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Neognathae Opisthokonta Primates Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms
G97531BE	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-4)Man(?1-	Mannose O-Mannose Polysialic acid	Linkage defined saccharide	836.29	Amniota Bifurcata Bilateria Chordata Colubroidea Craniata Daboia russelii russelii Daboia russelii Daboia Deuterostomia Dipnotetrapodomorpha Episquamata Eukaryota Eumetazoa Euteleostomi Gnathostomata Lepidosauria Metazoa Opisthokonta Sarcopterygii Sauria Sauropsida Serpentes Squamata Teleostomi Tetrapoda Toxicofera Unidentata Vertebrata Viperidae Viperinae cellular organisms
G98140IX	GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-4)][Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan core basic	Linkage defined saccharide	1316.49	Amniota Archelosauria Archosauria Aves Bilateria Boreoeutheria Catarrhini Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Neognathae Opisthokonta Phasianidae Phasianinae Primates Rattus norvegicus Rattus Rodentia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms
G98544DH	Neu5Ac(a2-8)Neu5Ac(a2-3)Gal(b1-4)Glc(b1-	9-O-Acetyl GD3 GD3	Linkage defined saccharide	924.31	Actinopteri Actinopterygii Amniota Amphibia Anura Archelosauria Archosauria Artiodactyla Aves Bacteria Balaenoptera edeni Balaenoptera Balaenopteridae Batoidea Batrachia Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Campylobacter jejuni Campylobacter Campylobacteraceae Campylobacterales Campylobacterota Canidae Caniformia Canis lupus familiaris Canis lupus Canis Carassius auratus Carassius Carnivora Catarrhini Cavia porcellus Cavia Caviidae Cercopithecidae Cercopithecinae Cercopithecoidea Cetacea Chinchilla Chinchillidae Chondrichthyes Chordata Clupeocephala Coelurosauria Craniata Cricetidae Cricetinae Cyprinidae Cypriniformes Cyprininae Cypriniphysae Cyprinoidei Deuterostomia Dinosauria Dipnotetrapodomorpha Discopyge tschudii Discopyge Elasmobranchii Epsilonproteobacteria Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Felidae Feliformia Felinae Felis catus Felis Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Hystricomorpha Lagomorpha Laurasiatheria Leporidae Leucoraja erinacea Leucoraja Macaca fuscata Macaca mulatta Macaca Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Mysticeti Narcinidae Neognathae Neopterygii Opisthokonta Oryctolagus cuniculus Oryctolagus Ostariophysi Osteoglossocephalai Otomorpha Otophysi Papio hamadryas Papio Pecora Phasianidae Phasianinae Pipidae Pipoidea Primates Rajidae Rajiformes Rattus Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Theropoda Torpedinidae Torpediniformes Torpedo marmorata Torpedo Vertebrata Whippomorpha Xenopodinae Xenopus cellular organisms
G98602UY	GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-4)][Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan hybrid	Linkage defined saccharide	1640.59	Amniota Archelosauria Archosauria Artiodactyla Aves Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Laurasiatheria Mammalia Metazoa Neognathae Opisthokonta Pecora Phasianidae Phasianinae Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms unclassified entries unclassified sequences unidentified
G98855LF	GlcNAc(b1-2)Man(a1-3)[GalNAc(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	LDN LacdiNAc N-Glycan complex	Linkage defined saccharide	1665.62	Amniota Bifurcata Bilateria Bothrops moojeni Bothrops Chordata Colubroidea Craniata Crotalinae Deuterostomia Dipnotetrapodomorpha Episquamata Eukaryota Eumetazoa Euteleostomi Gnathostomata Lepidosauria Metazoa Opisthokonta Sarcopterygii Sauria Sauropsida Serpentes Squamata Teleostomi Tetrapoda Toxicofera Unidentata Vertebrata Viperidae cellular organisms
G98894JN	Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-4)]Man(a1-3)[Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	GM4 N-Glycan complex Type 1 LN	Linkage defined saccharide	3681.3	Amniota Bifurcata Bilateria Calloselasma rhodostoma Calloselasma Chordata Colubroidea Craniata Crotalinae Deuterostomia Dipnotetrapodomorpha Episquamata Eukaryota Eumetazoa Euteleostomi Gnathostomata Lepidosauria Metazoa Opisthokonta Sarcopterygii Sauria Sauropsida Serpentes Squamata Teleostomi Tetrapoda Toxicofera Unidentata Vertebrata Viperidae cellular organisms
G99182WQ	Gal(b1-4)GlcNAc(b1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	I' antigen N-Glycan complex Type 2 LN2	Linkage defined saccharide	2370.86	Amniota Archelosauria Archosauria Aves Bilateria Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Metazoa Neognathae Opisthokonta Phasianidae Phasianinae Sarcopterygii Sauria Saurischia Sauropsida Teleostomi Tetrapoda Theropoda Vertebrata cellular organisms
G99524KA	Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-3)[Neu5Ac(a2-3)Gal(b1-3)GalNAc(b1-4)]Gal(b1-4)Glc(b1-	GQ1c	Linkage defined saccharide	1871.63	Acanthomorphata Actinopteri Actinopterygii Amniota Archelosauria Archosauria Aves Bilateria Boreoeutheria Catarrhini Chordata Cichlidae Cichliformes Cichlomorphae Clupeocephala Coelurosauria Craniata Ctenosquamata Deuterostomia Dinosauria Dipnotetrapodomorpha Euacanthomorphacea Euarchontoglires Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Neognathae Neopterygii Neoteleostei Opisthokonta Osteoglossocephalai Ovalentaria Percomorphaceae Phasianidae Phasianinae Primates Rattus Rodentia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostei Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms
G99927HL	GlcNAc(b1-4)Man(a1-3)[GlcNAc(b1-4)][GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan complex	Linkage defined saccharide	1722.65	Amniota Archelosauria Archosauria Aves Bilateria Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Metazoa Neognathae Opisthokonta Phasianidae Phasianinae Sarcopterygii Sauria Saurischia Sauropsida Teleostomi Tetrapoda Theropoda Vertebrata cellular organisms
G00032MO	Gal(b1-3)GalNAc	O-Glycan core 1 fuzzy	Linkage defined saccharide	383.14	Amniota Archelosauria Archosauria Aves Bacteria Bilateria Boreoeutheria Campylobacter coli Campylobacter Campylobacteraceae Campylobacterales Campylobacterota Catarrhini Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Enterobacterales Enterobacteriaceae Epsilonproteobacteria Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Gammaproteobacteria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Neognathae Opisthokonta Phasianidae Phasianinae Primates Pseudomonadota Sarcopterygii Sauria Saurischia Sauropsida Shigella dysenteriae Shigella Simiiformes Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms
G09652ZI	GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-4)][GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2087.78	Amniota Archelosauria Archosauria Aves Bilateria Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Metazoa Neognathae Opisthokonta Phasianidae Phasianinae Sarcopterygii Sauria Saurischia Sauropsida Teleostomi Tetrapoda Theropoda Vertebrata cellular organisms
G17100OM	Neu5Ac(a2-?)Gal(b1-?)GalNAc(b1-?)Gal(a1-?)Gal(b1-?)Glc(b1-	Cytolipin R GL-7 globoseries ganglioside Mono-sulfated globopentaosylceramide SSEA-4	Linkage defined saccharide	1160.4	Amniota Archelosauria Archosauria Aves Bilateria Boreoeutheria Catarrhini Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Neognathae Opisthokonta Phasianidae Phasianinae Primates Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms
G24137GH	Neu5Ac(a2-3)Gal(b1-3)[Gal(a1-4)Gal(b1-4)Gal(b1-4)GlcNAc(b1-6)]GalNAc(a1-3)GalNAc-ol	GM4 N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1568.57	Amniota Apodidae Apodiformes Apodinae Archelosauria Archosauria Aves Bilateria Caprimulgimorphae Chordata Coelurosauria Collocalia Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Hirundinidae Metazoa Neognathae Opisthokonta Passeriformes Phasianidae Phasianinae Sarcopterygii Sauria Saurischia Sauropsida Sylvioidea Teleostomi Tetrapoda Theropoda Vertebrata cellular organisms
G25627PL	GalNAc6S(b1-4)GlcA(b1-3)GalNAc6S		Linkage defined saccharide	760.12	Amniota Archelosauria Archosauria Aves Bilateria Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Metazoa Neognathae Opisthokonta Phasianidae Phasianinae Sarcopterygii Sauria Saurischia Sauropsida Teleostomi Tetrapoda Theropoda Vertebrata cellular organisms
G34996GA	Man(a1-3)[Neu5Ac(a2-3)Gal(b1-3)GlcNAc(b1-2/6)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc-ol	GM4 Type 1 LN	Linkage defined saccharide	1714.63	Amniota Bifurcata Bilateria Calloselasma rhodostoma Calloselasma Chordata Colubroidea Craniata Crotalinae Deuterostomia Dipnotetrapodomorpha Episquamata Eukaryota Eumetazoa Euteleostomi Gnathostomata Lepidosauria Metazoa Opisthokonta Sarcopterygii Sauria Sauropsida Serpentes Squamata Teleostomi Tetrapoda Toxicofera Unidentata Vertebrata Viperidae cellular organisms