GlyCosmos Portal

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GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	August 18, 2024

Displaying entries **26 - 50** of **276** in total

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GlyTouCan ID	IUPAC Condensed	Motifs	Subsumption Level	Monoisotopic Mass	Organisms ▲
G72627IU	Glc(a1-3)Glc(a1-3)Glc		Linkage defined saccharide	504.17	Agaricomycetes incertae sedis Agaricomycetes Agaricomycotina Ascomycota Bacilli Bacillota Bacteria Basidiomycota Dikarya Enterobacterales Enterobacteriaceae Eukaryota Fungi Gammaproteobacteria Lactobacillaceae Lactobacillales Laetiporaceae Laetiporus sulphureus Laetiporus Leuconostoc mesenteroides Leuconostoc Opisthokonta Polyporales Pseudomonadota Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales Saccharomycetes Saccharomycotina Shigella flexneri Shigella Terrabacteria group cellular organisms saccharomyceta
G66305IS		Glucosylceramide			Agaricomycetes Agaricomycetidae Agaricomycotina Ascomycota Basidiomycota Boletaceae Boletales Boletineae Boletoideae Dikarya Eukaryota Fungi Opisthokonta Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales Saccharomycetes Saccharomycotina Tylopilus felleus Tylopilus cellular organisms saccharomyceta
G30409YH	Glc(b1-3)Glc(b1-3)Glc		Linkage defined saccharide	504.17	Agaricomycotina Alphaproteobacteria Ascomycota Bacteria Basidiomycota Bradyrhizobium japonicum Bradyrhizobium Cutaneotrichosporon cutaneum Cutaneotrichosporon Dikarya Eukaryota Fungi Hyphomicrobiales Nitrobacteraceae Opisthokonta Pseudomonadota Rhizobiaceae Rhodophyta Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales Saccharomycetes Saccharomycotina Sinorhizobium fredii group Sinorhizobium fredii Sinorhizobium/Ensifer group Sinorhizobium Tremellomycetes Trichosporonaceae Trichosporonales cellular organisms saccharomyceta
G03895OL		Mannose O-Mannose			Agaricomycotina Ascomycota Basidiomycota Cutaneotrichosporon cutaneum Cutaneotrichosporon Dikarya Eukaryota Fungi Opisthokonta Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales Saccharomycetes Saccharomycotina Tremellomycetes Trichosporonaceae Trichosporonales cellular organisms saccharomyceta
G87997PG	?-Gal?(?1-4)Gal(b1-4)Glc		Linkage defined saccharide	504.17	Agaricomycotina Ascomycota Basidiomycota Dikarya Eukaryota Fungi Opisthokonta Papiliotrema laurentii Papiliotrema Rhynchogastremaceae Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales Saccharomycetes Saccharomycotina Tremellales Tremellomycetes cellular organisms saccharomyceta
G21939WM		Glucosylceramide			Alcaligenaceae Alcaligenes faecalis Alcaligenes Amniota Ascomycota Bacteria Betaproteobacteria Bilateria Boreoeutheria Burkholderiales Chordata Craniata Deuterostomia Dikarya Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Fungi Glires Gnathostomata Hypocreaceae Hypocreales Hypocreomycetidae Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Pezizomycotina Pseudomonadota Rattus Rodentia Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales Saccharomycetes Saccharomycotina Sarcopterygii Sordariomycetes Teleostomi Tetrapoda Theria Trichoderma longibrachiatum Trichoderma Vertebrata cellular organisms leotiomyceta saccharomyceta sordariomyceta
G61491DK	Man(a1-	Mannose O-Mannose			Alphaproteobacteria Alteromonadales Amniota Andropogoneae Andropogonodae Ascomycota Aspergillaceae Aspergillus niger Aspergillus subgen. Circumdati Aspergillus Bacteria Bilateria Boreoeutheria Caulobacter vibrioides CB15 Caulobacter vibrioides Caulobacter Caulobacteraceae Caulobacterales Chordata Craniata Deuterostomia Dikarya Dipnotetrapodomorpha Embryophyta Enterobacterales Enterobacteriaceae Escherichia coli O55 Escherichia coli Escherichia Euarchontoglires Eukaryota Eumetazoa Euphyllophyta Eurotiales Eurotiomycetes Eurotiomycetidae Euteleostomi Eutheria Fungi Gammaproteobacteria Glires Gnathostomata Gunneridae Liliopsida Magnoliopsida Mammalia Mesangiospermae Metazoa Muridae Murinae Muroidea Myomorpha Nicotiana Nicotianeae Nicotianoideae Opisthokonta PACMAD clade Panicoideae Pentapetalae Petrosaviidae Pezizomycotina Poaceae Poales Pseudoalteromonadaceae Pseudoalteromonas tetraodonis Pseudoalteromonas Pseudomonadota Rattus Rodentia Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales Saccharomycetes Saccharomycotina Sarcopterygii Shigella Solanaceae Solanales Spermatophyta Streptophyta Streptophytina Teleostomi Tetrapoda Theria Tracheophyta Tripsacinae Vertebrata Viridiplantae Zea mays subsp. mays Zea mays Zea asterids cellular organisms commelinids eudicotyledons lamiids leotiomyceta saccharomyceta
G25248YP					Alphaproteobacteria Ascomycota Bacteria Dikarya Eukaryota Fungi Hyphomicrobiales Opisthokonta Pseudomonadota Rhizobiaceae Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales Saccharomycetes Saccharomycotina Sinorhizobium meliloti Sinorhizobium/Ensifer group Sinorhizobium cellular organisms saccharomyceta
G57930PW	Glc(a1-2)Glc(a1-3)Glc(a1-3)Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan high mannose	Linkage defined saccharide	2368.8	Alpharhabdovirinae Amniota Ascomycota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dikarya Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Fungi Glires Gnathostomata Haplorrhini Haploviricotina Hominidae Homininae Hominoidea Homo sapiens Homo Lyssavirus rabies Lyssavirus Mammalia Metazoa Monjiviricetes Mononegavirales Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Negarnaviricota Opisthokonta Orthornavirae Primates Rhabdoviridae Riboviria Rodentia Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales Saccharomycetes Saccharomycotina Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms saccharomyceta
G39578OU	Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(a1-	N-Glycan high mannose	Linkage defined saccharide	1882.64	Alpharhabdovirinae Ascomycota Dikarya Eukaryota Fungi Haploviricotina Monjiviricetes Mononegavirales Negarnaviricota Opisthokonta Orthornavirae Rhabdoviridae Riboviria Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales Saccharomycetes Saccharomycotina Vesicular stomatitis virus Vesiculovirus Viruses cellular organisms saccharomyceta unclassified Vesiculovirus
G63560NE	Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc		Linkage defined saccharide	707.25	Alphavirus Alsuviricetes Amniota Artiodactyla Ascomycota Bilateria Boreoeutheria Bovidae Caprinae Carnivora Chordata Craniata Deuterostomia Dikarya Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Felidae Feliformia Felinae Felis catus Felis Fungi Gnathostomata Kitrinoviricota Laurasiatheria Mammalia Martellivirales Metazoa Opisthokonta Orthornavirae Ovis aries Ovis Pecora Riboviria Ruminantia Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales Saccharomycetes Saccharomycotina Sarcopterygii Semliki Forest virus Teleostomi Tetrapoda Theria Togaviridae Vertebrata Viruses cellular organisms saccharomyceta
G41824PC	Glc(a1-2)Glc(a1-3)Glc(a1-3)Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan core basic	Linkage defined saccharide	1720.59	Alveolata Amniota Ascomycota Bilateria Boreoeutheria Chordata Ciliophora Craniata Deuterostomia Dikarya Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Fungi Glires Gnathostomata Hymenostomatida Intramacronucleata Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Oligohymenophorea Opisthokonta Rodentia Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales Saccharomycetes Saccharomycotina Sar Sarcopterygii Teleostomi Tetrahymena pyriformis Tetrahymena Tetrahymenidae Tetrahymenina Tetrapoda Theria Vertebrata cellular organisms saccharomyceta
G67828VR	Man(b1-4)GlcNAc(b1-4)GlcNAc		Linkage defined saccharide	586.22	Amniota Amphiesmenoptera Amphipyrinae Arthropoda Artiodactyla Ascomycota Bilateria Boreoeutheria Bovidae CS clade Capra hircus Capra Caprinae Chlamydomonadales Chlorophyceae Chlorophyta Chordata Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dicondylia Dikarya Dipnotetrapodomorpha Ditrysia Ecdysozoa Endopterygota Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Fungi Glires Glossata Gnathostomata Heteroneura Hexapoda Insecta Laurasiatheria Lepidoptera Mammalia Mandibulata Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neolepidoptera Neoptera Noctuidae Noctuoidea Obtectomera Opisthokonta Panarthropoda Pancrustacea Pecora Protostomia Pterygota Rattus Rodentia Ruminantia Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales Saccharomycetes Saccharomycotina Sarcopterygii Spodoptera frugiperda Spodoptera Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata Viridiplantae Volvocaceae Volvox carteri Volvox cellular organisms core chlorophytes saccharomyceta
G24934QN					Amniota Andropogoneae Andropogonodae Archelosauria Archosauria Ascomycota Aves Bilateria Chordata Coelurosauria Columba Columbidae Columbiformes Craniata Deuterostomia Dikarya Dinosauria Dipnotetrapodomorpha Embryophyta Eukaryota Eumetazoa Euphyllophyta Euteleostomi Fungi Gnathostomata Gunneridae Liliopsida Magnoliopsida Mesangiospermae Metazoa Neognathae Nicotiana tabacum Nicotiana Nicotianeae Nicotianoideae Opisthokonta PACMAD clade Panicoideae Pentapetalae Petrosaviidae Poaceae Poales Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales Saccharomycetes Saccharomycotina Sarcopterygii Sauria Saurischia Sauropsida Solanaceae Solanales Spermatophyta Streptophyta Streptophytina Teleostomi Tetrapoda Theropoda Tracheophyta Tripsacinae Vertebrata Viridiplantae Zea mays Zea asterids cellular organisms commelinids eudicotyledons lamiids saccharomyceta
G15407YE	Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-		Linkage defined saccharide	586.22	Amniota Archelosauria Archosauria Artiodactyla Ascomycota Aves Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Coelurosauria Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dikarya Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Fungi Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Neognathae Opisthokonta Pecora Phasianidae Phasianinae Primates Rattus Rodentia Ruminantia Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales Saccharomycetes Saccharomycotina Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms saccharomyceta
G19958IL	Glc(a1-3)Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan high mannose	Linkage defined saccharide	2044.7	Amniota Archelosauria Archosauria Ascomycota Aves Bilateria Boreoeutheria Catarrhini Chordata Coelurosauria Coturnix japonica Coturnix Craniata Deuterostomia Dikarya Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Fungi Galliformes Galloanserae Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Neognathae Opisthokonta Perdicinae Phasianidae Primates Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales Saccharomycetes Saccharomycotina Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms saccharomyceta
G65996GC	Man(b1-4)GlcNAc(b1-4)GlcNAc(a1-		Linkage defined saccharide	586.22	Amniota Archelosauria Archosauria Ascomycota Aves Bilateria Chordata Coelurosauria Craniata Deuterostomia Dikarya Dinosauria Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Fungi Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Metazoa Neognathae Opisthokonta Phasianidae Phasianinae Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales Saccharomycetes Saccharomycotina Sarcopterygii Sauria Saurischia Sauropsida Teleostomi Tetrapoda Theropoda Vertebrata cellular organisms saccharomyceta
G46565HP	Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc		Linkage defined saccharide	1517.51	Amniota Arthropoda Artiodactyla Ascomycota Bilateria Blaberoidea Blattella germanica Blattella Blattellidae Blattodea Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deuterostomia Dicondylia Dictyoptera Dikarya Dipnotetrapodomorpha Ecdysozoa Embryophyta Euarchontoglires Eukaryota Eumetazoa Euphyllophyta Euteleostomi Eutheria Fungi incertae sedis Fungi Glires Gnathostomata Gunneridae Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Laurasiatheria Magnoliopsida Mammalia Mandibulata Mesangiospermae Metazoa Mucorales Mucorineae Mucoromycetes Mucoromycota Mucoromycotina Muridae Murinae Muroidea Myomorpha Neoptera Opisthokonta Panarthropoda Pancrustacea Pecora Pentapetalae Polyneoptera Primates Protostomia Pterygota Rattus Rhizopodaceae Rhizopus niveus Rhizopus Rodentia Ruminantia Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales Saccharomycetes Saccharomycotina Sarcopterygii Simiiformes Solanaceae Solanales Solaneae Solanoideae Solanum lycopersicum Solanum subgen. Lycopersicon Solanum Spermatophyta Streptophyta Streptophytina Teleostomi Tetrapoda Theria Tracheophyta Vertebrata Viridiplantae asterids cellular organisms eudicotyledons lamiids saccharomyceta
G18546YL	Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc		Linkage defined saccharide	1679.57	Amniota Arthropoda Artiodactyla Ascomycota Bilateria Blattodea Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deuterostomia Dicondylia Dictyoptera Dikarya Dipnotetrapodomorpha Ecdysozoa Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Fungi Glires Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Laurasiatheria Mammalia Mandibulata Metazoa Muridae Murinae Muroidea Myomorpha Neoptera Opisthokonta Panarthropoda Pancrustacea Pecora Polyneoptera Primates Protostomia Pterygota Rattus Rodentia Ruminantia Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales Saccharomycetes Saccharomycotina Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms saccharomyceta
G83565SN	Glc(a1-3)Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc		Linkage defined saccharide	1841.62	Amniota Arthropoda Ascomycota Bilateria Blaberoidea Blattella germanica Blattella Blattellidae Blattodea Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dicondylia Dictyoptera Dikarya Dipnotetrapodomorpha Ecdysozoa Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Fungi Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Mammalia Mandibulata Metazoa Neoptera Opisthokonta Panarthropoda Pancrustacea Polyneoptera Primates Protostomia Pterygota Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales Saccharomycetes Saccharomycotina Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms saccharomyceta
G16794FO	Man(a1-3)Man		Linkage defined saccharide	342.12	Amniota Artiodactyla Ascomycota Bilateria Boreoeutheria CUG-Ala clade Chordata Craniata Deuterostomia Dikarya Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Fungi Gnathostomata Laurasiatheria Mammalia Metazoa Nakazawaea holstii Nakazawaea Opisthokonta Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales incertae sedis Saccharomycetales Saccharomycetes Saccharomycotina Sarcopterygii Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms saccharomyceta
G26534TO	Glc(a1-3)Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan core basic	Linkage defined saccharide	1396.49	Amniota Artiodactyla Ascomycota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dikarya Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Fungi Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales Saccharomycetes Saccharomycotina Sarcopterygii Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms saccharomyceta
G25559RS	Man(a1-2)Man(a1-2)Man(a1-	Mannose O-Mannose	Linkage defined saccharide	504.17	Amniota Ascomycota Bilateria Boreoeutheria CUG-Ser1 clade Candida albicans Candida/Lodderomyces clade Candida Catarrhini Chordata Craniata Debaryomycetaceae Deuterostomia Dikarya Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Fungi Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Komagataella pastoris Komagataella Mammalia Metazoa Opisthokonta Phaffomycetaceae Primates Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales Saccharomycetes Saccharomycotina Sarcopterygii Schizosaccharomyces pombe Schizosaccharomyces Schizosaccharomycetaceae Schizosaccharomycetales Schizosaccharomycetes Simiiformes Taphrinomycotina Teleostomi Tetrapoda Theria Vertebrata cellular organisms saccharomyceta
G59092IU	Glc(a1-2)Glc(a1-3)Glc(a1-3)Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc		Linkage defined saccharide	2165.72	Amniota Ascomycota Bilateria Boreoeutheria Catarrhini Chordata Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dikarya Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Fungi Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muroidea Myomorpha Opisthokonta Primates Rodentia Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales Saccharomycetes Saccharomycotina Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms saccharomyceta
G52878KK	Man(a1-?)Man		Linkage defined saccharide	342.12	Amniota Ascomycota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dikarya Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Fungi Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Opisthokonta Primates Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales Saccharomycetes Saccharomycotina Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms saccharomyceta