GlyCosmos Portal

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GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	August 18, 2024

Displaying entries **8126 - 8150** of **9278** in total

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GlyTouCan ID	IUPAC Condensed	Motifs	Subsumption Level	Monoisotopic Mass	Organisms ▼
G78160OZ	GlcA(b1-3)GalNAc4S		Linkage defined saccharide	477.08	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Canidae Caniformia Canis lupus familiaris Canis lupus Canis Carnivora Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Lagomorpha Laurasiatheria Leporidae Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Oryctolagus cuniculus Oryctolagus Pecora Primates Rattus Rodentia Ruminantia Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G07982RO	GlcNAc(b1-4)GlcNAc(?1-		Linkage defined saccharide	424.17	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Brassicales Carica papaya Carica Caricaceae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Embryophyta Euarchontoglires Eukaryota Eumetazoa Euphyllophyta Euteleostomi Eutheria Gnathostomata Gunneridae Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Magnoliopsida Mammalia Mesangiospermae Metazoa Opisthokonta Pecora Pentapetalae Primates Ruminantia Sarcopterygii Simiiformes Spermatophyta Streptophyta Streptophytina Teleostomi Tetrapoda Theria Tracheophyta Vertebrata Viridiplantae cellular organisms eudicotyledons malvids rosids
G97179UV	Glc(a1-3)Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-3)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan core basic	Linkage defined saccharide	1558.54	Amniota Artiodactyla Bilateria Blastocrithidia Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Discoba Euglenozoa Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Herpetomonas samuelpessoai Herpetomonas Kinetoplastea Laurasiatheria Leishmania adleri Leishmania mexicana species complex Leishmania mexicana Leishmania Leishmania Leishmaniinae Leptomonas samueli Leptomonas Mammalia Metakinetoplastina Metazoa Opisthokonta Sarcopterygii Schizotrypanum Strigomonadinae Strigomonas culicis Strigomonas Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Trypanosoma conorhini Trypanosoma dionisii Trypanosoma with unspecified subgenus Trypanosoma Trypanosomatida Trypanosomatidae Vertebrata cellular organisms lizard Leishmania
G44770ZI	GalNAc(b1-4)GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	LDN LacdiNAc N-Glycan complex	Linkage defined saccharide	1665.62	Amniota Artiodactyla Bifurcata Bilateria Boreoeutheria Bothrops moojeni Bothrops Catarrhini Chordata Colubroidea Craniata Crotalinae Deuterostomia Dipnotetrapodomorpha Episquamata Equidae Equus caballus Equus Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Lepidosauria Mammalia Metazoa Opisthokonta Perissodactyla Primates Sarcopterygii Sauria Sauropsida Serpentes Simiiformes Squamata Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Toxicofera Unidentata Vertebrata Viperidae cellular organisms
G78342SP	GalNAc(b1-4)GlcNAc(b1-2)Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	LDN LacdiNAc N-Glycan core basic	Linkage defined saccharide	1462.54	Amniota Artiodactyla Bifurcata Bilateria Boreoeutheria Bothrops moojeni Bothrops Catarrhini Chordata Colubroidea Craniata Crotalinae Deuterostomia Dipnotetrapodomorpha Episquamata Equidae Equus caballus Equus Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Lepidosauria Mammalia Metazoa Opisthokonta Perissodactyla Primates Sarcopterygii Sauria Sauropsida Serpentes Simiiformes Squamata Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Toxicofera Unidentata Vertebrata Viperidae cellular organisms
G59624GO	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-4)Man	Polysialic acid	Linkage defined saccharide	836.29	Amniota Artiodactyla Bifurcata Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Colubroidea Craniata Daboia russelii russelii Daboia russelii Daboia Deuterostomia Dipnotetrapodomorpha Episquamata Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Lepidosauria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Sauria Sauropsida Serpentes Squamata Teleostomi Tetrapoda Theria Toxicofera Unidentata Vertebrata Viperidae Viperinae cellular organisms
G39852JB		N-acetyllactosamine Type 2 LN			Amniota Artiodactyla Bifurcata Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Colubroidea Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dipnotetrapodomorpha Elapidae Elapinae Episquamata Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Lepidosauria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Naja Opisthokonta Pecora Primates Rodentia Ruminantia Sarcopterygii Sauria Sauropsida Serpentes Simiiformes Squamata Teleostomi Tetrapoda Theria Toxicofera Unidentata Vertebrata cellular organisms
G11960UF	GalNAc(b1-4)GlcNAc(b1-2)Man(a1-3)[GalNAc(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	LDN LacdiNAc N-Glycan complex	Linkage defined saccharide	1722.65	Amniota Artiodactyla Bifurcata Bilateria Boreoeutheria Bos taurus Bos Bothrops moojeni Bothrops Bovidae Bovinae Catarrhini Chordata Colubroidea Craniata Crotalinae Deuterostomia Dipnotetrapodomorpha Episquamata Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Lepidosauria Mammalia Metazoa Opisthokonta Pecora Primates Ruminantia Sarcopterygii Sauria Sauropsida Serpentes Simiiformes Squamata Teleostomi Tetrapoda Theria Toxicofera Unidentata Vertebrata Viperidae cellular organisms
G60653VD	GalNAc(b1-4)GlcNAc(b1-2)Man(a1-3)[GalNAc(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	LDN LacdiNAc N-Glycan complex	Linkage defined saccharide	1868.7	Amniota Artiodactyla Bifurcata Bilateria Boreoeutheria Bos taurus Bos Bothrops moojeni Bothrops Bovidae Bovinae Catarrhini Chordata Colubroidea Craniata Cricetidae Cricetinae Crotalinae Deuterostomia Digenea Dipnotetrapodomorpha Episquamata Equidae Equus caballus Equus Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Lepidosauria Lophotrochozoa Mammalia Mesocricetus auratus Mesocricetus Metazoa Muroidea Myomorpha Opisthokonta Pecora Perissodactyla Platyhelminthes Primates Protostomia Rodentia Ruminantia Sarcopterygii Sauria Sauropsida Schistosoma mansoni Schistosoma Schistosomatidae Schistosomatoidea Serpentes Simiiformes Spiralia Squamata Strigeidida Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Toxicofera Trematoda Unidentata Vertebrata Viperidae cellular organisms
G93320VA	Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(a1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	N-Glycan complex N-acetyllactosamine Type 2 LN a-Gal antigen	Linkage defined saccharide	1948.7	Amniota Artiodactyla Batoidea Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chondrichthyes Chordata Craniata Deuterostomia Dipnotetrapodomorpha Elasmobranchii Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Pecora Rodentia Ruminantia Sarcopterygii Teleostomi Tetrapoda Tetronarce californica Tetronarce Theria Torpedinidae Torpediniformes Vertebrata cellular organisms
G39992DR	Gal(a1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(a1-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	N-Glycan complex N-acetyllactosamine Type 2 LN a-Gal antigen	Linkage defined saccharide	2110.76	Amniota Artiodactyla Batoidea Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chondrichthyes Chordata Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dipnotetrapodomorpha Elasmobranchii Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Pecora Rodentia Ruminantia Sarcopterygii Teleostomi Tetrapoda Tetronarce californica Tetronarce Theria Torpedinidae Torpediniformes Vertebrata cellular organisms
G70059SD	GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1624.6	Amniota Artiodactyla Batoidea Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chondrichthyes Chordata Craniata Deuterostomia Dipnotetrapodomorpha Elasmobranchii Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Pecora Primates Rodentia Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Tetronarce californica Tetronarce Theria Torpedinidae Torpediniformes Vertebrata cellular organisms
G86102WJ	GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1478.54	Amniota Artiodactyla Batoidea Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chondrichthyes Chordata Craniata Deuterostomia Dipnotetrapodomorpha Elasmobranchii Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Pecora Primates Rodentia Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Tetronarce californica Tetronarce Theria Torpedinidae Torpediniformes Vertebrata cellular organisms
G99129GB	Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1478.54	Amniota Artiodactyla Batoidea Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chondrichthyes Chordata Craniata Deuterostomia Dipnotetrapodomorpha Elasmobranchii Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Pecora Primates Rodentia Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Tetronarce californica Tetronarce Theria Torpedinidae Torpediniformes Vertebrata cellular organisms
G84678BK	GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan complex	Linkage defined saccharide	1316.49	Amniota Artiodactyla Batoidea Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chondrichthyes Chordata Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dipnotetrapodomorpha Elasmobranchii Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Pecora Primates Rodentia Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Tetronarce californica Tetronarce Theria Torpedinidae Torpediniformes Vertebrata cellular organisms
G80343LW	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2005.72	Amniota Artiodactyla Batoidea Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Carnivora Catarrhini Chondrichthyes Chordata Craniata Deuterostomia Dipnotetrapodomorpha Elasmobranchii Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Felidae Feliformia Felinae Felis catus Felis Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Primates Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Tetronarce californica Tetronarce Theria Torpedinidae Torpediniformes Vertebrata cellular organisms
G88800UV	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2370.86	Amniota Artiodactyla Batoidea Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Carnivora Catarrhini Chondrichthyes Chordata Craniata Deuterostomia Dipnotetrapodomorpha Elasmobranchii Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Felidae Feliformia Felinae Felis catus Felis Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Pecora Primates Rodentia Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Tetronarce californica Tetronarce Theria Torpedinidae Torpediniformes Vertebrata cellular organisms
G69331UE			Linkage defined saccharide	1966.11	Amniota Artiodactyla Balaenoptera physalus Balaenoptera Balaenopteridae Bilateria Boreoeutheria Cetacea Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Mysticeti Opisthokonta Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata Whippomorpha cellular organisms
G22856AB	Gal(b1-4)GlcNAc(b1-3)[Gal(b1-4)GlcNAc(b1-6)]Gal	N-acetyllactosamine Type 2 LN	Linkage defined saccharide	910.33	Amniota Artiodactyla Bacteria Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Enterobacterales Enterobacteriaceae Escherichia coli Escherichia Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gammaproteobacteria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Opisthokonta Primates Pseudomonadota Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G00068MO	Fuc(a1-2)Gal(b1-	Galactosylceramide	Linkage defined saccharide	326.12	Amniota Artiodactyla Bacteria Bilateria Boreoeutheria Campylobacterales Campylobacterota Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Epsilonproteobacteria Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Helicobacter pylori Helicobacter Helicobacteraceae Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G03071KV	GalNAc(a1-3)[Fuc(a1-2)]Gal(b1-4)GlcNAc		Linkage defined saccharide	732.28	Amniota Artiodactyla Bacteria Bilateria Boreoeutheria Campylobacterales Campylobacterota Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Epsilonproteobacteria Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Helicobacter pylori Helicobacter Helicobacteraceae Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Primates Rattus Rodentia Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G06171MB	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-6)]Man	N-acetyllactosamine Type 2 LN	Linkage defined saccharide	910.33	Amniota Artiodactyla Bacteria Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Enterobacterales Enterobacteriaceae Escherichia coli Escherichia Eukaryota Eumetazoa Euteleostomi Eutheria Gammaproteobacteria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Pseudomonadota Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G32403EK	Gal(b1-4)GlcNAc(b1-6)Man	N-acetyllactosamine Type 2 LN	Linkage defined saccharide	545.2	Amniota Artiodactyla Bacteria Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Enterobacterales Enterobacteriaceae Escherichia coli Escherichia Eukaryota Eumetazoa Euteleostomi Eutheria Gammaproteobacteria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Pseudomonadota Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G62262YL	Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man	N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1599.57	Amniota Artiodactyla Bacteria Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Enterobacterales Enterobacteriaceae Escherichia coli Escherichia Eukaryota Eumetazoa Euteleostomi Eutheria Gammaproteobacteria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Pseudomonadota Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G99210HQ	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man	N-acetyllactosamine Type 2 LN	Linkage defined saccharide	910.33	Amniota Artiodactyla Bacteria Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Enterobacterales Enterobacteriaceae Escherichia coli Escherichia Eukaryota Eumetazoa Euteleostomi Eutheria Gammaproteobacteria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Pseudomonadota Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms