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GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	August 18, 2024

Displaying entries **126 - 150** of **393** in total

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GlyTouCan ID	IUPAC Condensed	Motifs	Subsumption Level	Monoisotopic Mass ▼	Organisms
G53539GK	Neu5Gc(a2-3)[GalNAc(b1-4)]GalNAc(b1-3)Gal(b1-4)Gal(b1-3)[Neu5Gc(a2-8)Neu5Gc(a2-6)]GalNAc(?1-	O-Glycan core 1 fuzzy	Linkage defined saccharide	1872.63	Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Deuterostomia Eukaryota Eumetazoa Euteleosteomorpha Euteleostomi Gnathostomata Metazoa Neopterygii Oncorhynchus mykiss Oncorhynchus Opisthokonta Osteoglossocephalai Protacanthopterygii Salmonidae Salmoniformes Salmoninae Salvelinus leucomaenis pluvius Salvelinus leucomaenis Salvelinus Teleostei Teleostomi Vertebrata cellular organisms
G54442GQ	Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-3)[Neu5Ac(a2-3)Gal(b1-3)GalNAc(b1-4)]Gal(b1-4)Glc(?1-	GM4	Linkage defined saccharide	1871.63	Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Eukaryota Eumetazoa Euteleostomi Gnathostomata Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Teleostei Teleostomi Vertebrata cellular organisms
G18625KA	Neu5Ac(a2-8)Neu5Ac(a2-3)[Neu5Ac(a2-8)Neu5Ac(a2-3)Gal(b1-3)GalNAc(b1-4)]Gal(b1-4)Glc(b1-	GQ1b	Linkage defined saccharide	1871.63	Acanthomorphata Actinopteri Actinopterygii African cichlids Amniota Archelosauria Archosauria Artiodactyla Aves Bacteria Balaenoptera acutorostrata Balaenoptera brydei Balaenoptera edeni Balaenoptera Balaenopteridae Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Campylobacter jejuni Campylobacter Campylobacteraceae Campylobacterales Campylobacterota Carassius auratus Carassius Carnivora Catarrhini Cetacea Channichthyidae Chinchilla Chinchillidae Chionodraco myersi Chionodraco Chordata Cichlidae Cichliformes Cichlomorphae Clupeocephala Coelurosauria Craniata Cricetidae Cricetinae Ctenosquamata Cyprinidae Cypriniformes Cyprininae Cypriniphysae Cyprinoidei Cyprinus carpio Cyprinus Delphinidae Delphinus delphis Delphinus Deuterostomia Dinosauria Dipnotetrapodomorpha Epsilonproteobacteria Euacanthomorphacea Euarchontoglires Eukaryota Eumetazoa Eupercaria Eurypterygia Euteleosteomorpha Euteleostomi Eutheria Felidae Feliformia Felinae Felis catus Felis Galliformes Galloanserae Gallus gallus Gallus Glires Globicephala macrorhynchus Globicephala Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Hystricomorpha Lagomorpha Laurasiatheria Leporidae Leuciscidae Leuciscinae Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Mysticeti Neognathae Neopterygii Neoteleostei Nototheniidae Notothenioidei Odontoceti Opisthokonta Orcinus orca Orcinus Oreochromini Oreochromis mossambicus Oreochromis Oryctolagus cuniculus Oryctolagus Ostariophysi Osteoglossocephalai Otomorpha Otophysi Ovalentaria Pecora Perciformes Percomorphaceae Phasianidae Phasianinae Phocoenidae Phocoenoides dalli Phocoenoides Phodopus sungorus Phodopus Physeter catodon Physeter Physeteridae Primates Pseudocrenilabrinae Rattus Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Scardinius erythrophthalmus Scardinius Simiiformes Stenella coeruleoalba Stenella Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Theropoda Trematomus lepidorhinus Trematomus Tursiops truncatus Tursiops Vertebrata Whippomorpha cellular organisms
G54434YJ	Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-3)[Neu5Ac(a2-3)Gal(b1-3)GalNAc(b1-4)]Gal(b1-4)Glc	GM4	Linkage defined saccharide	1871.63	Acanthomorphata Actinopteri Actinopterygii Amniota Archelosauria Archosauria Aves Bilateria Chordata Clupeocephala Coelurosauria Craniata Ctenosquamata Deuterostomia Dinosauria Dipnotetrapodomorpha Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Gadariae Gadidae Gadiformes Gadoidei Gadus chalcogrammus Gadus Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Metazoa Neognathae Neopterygii Neoteleostei Opisthokonta Osteoglossocephalai Paracanthopterygii Phasianidae Phasianinae Pollachius pollachius Pollachius Sarcopterygii Sauria Saurischia Sauropsida Teleostei Teleostomi Tetrapoda Theropoda Vertebrata Zeiogadaria cellular organisms
G99524KA	Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-3)[Neu5Ac(a2-3)Gal(b1-3)GalNAc(b1-4)]Gal(b1-4)Glc(b1-	GQ1c	Linkage defined saccharide	1871.63	Acanthomorphata Actinopteri Actinopterygii Amniota Archelosauria Archosauria Aves Bilateria Boreoeutheria Catarrhini Chordata Cichlidae Cichliformes Cichlomorphae Clupeocephala Coelurosauria Craniata Ctenosquamata Deuterostomia Dinosauria Dipnotetrapodomorpha Euacanthomorphacea Euarchontoglires Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Neognathae Neopterygii Neoteleostei Opisthokonta Osteoglossocephalai Ovalentaria Percomorphaceae Phasianidae Phasianinae Primates Rattus Rodentia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostei Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms
G71158TZ	Neu5Ac(a2-8)Neu5Ac(a2-3)[Neu5Ac(a2-8)Neu5Ac(a2-3)Gal(b1-3)GalNAc(b1-4)]Gal(b1-4)Glc	GM4	Linkage defined saccharide	1871.63	Acanthomorphata Actinopteri Actinopterygii Amniota Archelosauria Archosauria Aves Bilateria Chordata Clupeocephala Coelurosauria Craniata Ctenosquamata Deuterostomia Dinosauria Dipnotetrapodomorpha Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Gadariae Gadidae Gadiformes Gadoidei Gadus chalcogrammus Gadus Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Metazoa Neognathae Neopterygii Neoteleostei Opisthokonta Osteoglossocephalai Paracanthopterygii Phasianidae Phasianinae Pollachius pollachius Pollachius Sarcopterygii Sauria Saurischia Sauropsida Teleostei Teleostomi Tetrapoda Theropoda Vertebrata Zeiogadaria cellular organisms
G59965VP	Gal(a1-3)[GalNAc(b1-4)GlcNAc(b1-3)Gal(b1-4)]Gal(b1-4)GlcNAc(b1-?)Man(a1-3)Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	LDN LacdiNAc N-acetyllactosamine Type 2 LN a-Gal antigen	Linkage defined saccharide	1843.67	Acanthomorphata Actinopteri Actinopterygii Atherinomorphae Bilateria Chordata Clupeocephala Craniata Ctenosquamata Cyprinodontiformes Cyprinodontoidei Deuterostomia Euacanthomorphacea Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Fundulidae Fundulus heteroclitus Fundulus Gnathostomata Metazoa Neopterygii Neoteleostei Opisthokonta Osteoglossocephalai Ovalentaria Percomorphaceae Teleostei Teleostomi Vertebrata cellular organisms
G95920TH	Gal(a1-3)[GalNAc(b1-4)GlcNAc(b1-3)Gal(b1-4)]Gal(b1-4)GlcNAc(b1-?)Man(a1-3/6)Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	LDN LacdiNAc N-acetyllactosamine Type 2 LN a-Gal antigen	Linkage defined saccharide	1843.67	Acanthomorphata Actinopteri Actinopterygii Atherinomorphae Bilateria Chordata Clupeocephala Craniata Ctenosquamata Cyprinodontiformes Cyprinodontoidei Deuterostomia Euacanthomorphacea Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Fundulidae Fundulus heteroclitus Fundulus Gnathostomata Metazoa Neopterygii Neoteleostei Opisthokonta Osteoglossocephalai Ovalentaria Percomorphaceae Teleostei Teleostomi Vertebrata cellular organisms
G10721QW	Neu5Gc(a2-3)[GalNAc(b1-4)]GalNAc(b1-4)Gal(b1-3)Gal(b1-3)[Neu5Ac(a2-8)Neu5Ac(a2-6)]GalNAc(a1-	O-Glycan core 1 Sialyl Tn antigen T antigen TF antigen	Linkage defined saccharide	1840.64	Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Deuterostomia Eukaryota Eumetazoa Euteleosteomorpha Euteleostomi Gnathostomata Metazoa Neopterygii Oncorhynchus mykiss Oncorhynchus Opisthokonta Osteoglossocephalai Protacanthopterygii Salmonidae Salmoniformes Salmoninae Teleostei Teleostomi Vertebrata cellular organisms
G21508HF	Gal(b1-4)GlcNAc(b1-2)Man(a1-?)[GlcNAc(b1-2)Man(a1-?)][GlcNAc(b1-4)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1827.68	Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Eukaryota Eumetazoa Euteleostomi Gnathostomata Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Teleostei Teleostomi Vertebrata cellular organisms
G33718MC	Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-?)[GlcNAc(b1-2)Man(a1-?)][GlcNAc(b1-4)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	Lewis x Lex N-Glycan complex N-acetyllactosamine SSEA-1 Type 2 LN	Linkage defined saccharide	1827.68	Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Eukaryota Eumetazoa Euteleostomi Gnathostomata Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Teleostei Teleostomi Vertebrata cellular organisms
G49664YG	GalNAc(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	LDN LacdiNAc N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1827.68	Actinopteri Actinopterygii Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Pecora Primates Ruminantia Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G69943PA	Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-?)[GalNAc(b1-4)GlcNAc(b1-2)Man(a1-?)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	LDN LacdiNAc Lewis x Lex N-Glycan complex N-acetyllactosamine SSEA-1 Type 2 LN	Linkage defined saccharide	1827.68	Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Eukaryota Eumetazoa Euteleostomi Gnathostomata Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Teleostei Teleostomi Vertebrata cellular organisms
G55717XU	Fuc(a1-3)GalNAc(b1-3)Gal(b1-4)Gal(b1-3)[Neu5Gc(a2-8)Neu5Gc(a2-8)Neu5Gc(a2-6)]GalNAc-ol		Linkage defined saccharide	1817.62	Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Deuterostomia Eukaryota Eumetazoa Euteleosteomorpha Euteleostomi Gnathostomata Metazoa Neopterygii Oncorhynchus keta Oncorhynchus Opisthokonta Osteoglossocephalai Protacanthopterygii Salmonidae Salmoniformes Salmoninae Teleostei Teleostomi Vertebrata cellular organisms
G00998NI	Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1786.65	Actinopteri Actinopterygii Alphainfluenzavirus influenzae Alphainfluenzavirus Amniota Articulavirales Bilateria Boreoeutheria Catarrhini Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata H7N7 subtype Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Influenza A virus (A/FPV/Dutch/1927(H7N7)) Influenza A virus Insthoviricetes Mammalia Metazoa Negarnaviricota Neopterygii Opisthokonta Orthomyxoviridae Orthornavirae Ostariophysi Osteoglossocephalai Otomorpha Otophysi Polyploviricotina Primates Riboviria Sarcopterygii Simiiformes Teleostei Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms unclassified Orthomyxoviridae unclassified entries unclassified sequences unidentified influenza virus unidentified
G86282DQ	Gal(b1-4)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-?)[GlcNAc(b1-2)Man(a1-?)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	Lewis x Lex N-Glycan complex N-acetyllactosamine SSEA-1 Type 2 LN	Linkage defined saccharide	1786.65	Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Eukaryota Eumetazoa Euteleostomi Gnathostomata Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Teleostei Teleostomi Vertebrata cellular organisms
G82560WA	Neu5Gc(a2-3)GalNAc(b1-4)GlcNAc(b1-2)Man(a1-?)[Man(a1-?)Man(a1-?)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	LDN LacdiNAc N-Glycan hybrid	Linkage defined saccharide	1785.63	Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Eukaryota Eumetazoa Euteleostomi Gnathostomata Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Teleostei Teleostomi Vertebrata cellular organisms
G92284FK	Gal(b1-4)Gal(b1-3)[Neu5Gc(a2-8)Neu5Gc(a2-8)Neu5Gc(a2-8)Neu5Gc(a2-6)]GalNAc-ol		Linkage defined saccharide	1775.57	Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Deuterostomia Eukaryota Eumetazoa Euteleosteomorpha Euteleostomi Gnathostomata Metazoa Neopterygii Oncorhynchus masou ishikawae Oncorhynchus masou Oncorhynchus mykiss Oncorhynchus Opisthokonta Osteoglossocephalai Protacanthopterygii Salmonidae Salmoniformes Salmoninae Salvelinus leucomaenis japonicus Salvelinus leucomaenis pluvius Salvelinus leucomaenis Salvelinus Teleostei Teleostomi Vertebrata cellular organisms
G37165RI	Gal(b1-4)Gal(b1-3)[Neu5Gc(a2-8)Neu5Gc(a2-8)Neu5Gc(a2-8)Neu5Gc(a2-6)]GalNAc(?1-	O-Glycan core 1 fuzzy	Linkage defined saccharide	1773.56	Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Deuterostomia Eukaryota Eumetazoa Euteleosteomorpha Euteleostomi Gnathostomata Metazoa Neopterygii Opisthokonta Osteoglossocephalai Protacanthopterygii Salmonidae Salmoniformes Salmoninae Salvelinus leucomaenis pluvius Salvelinus leucomaenis Salvelinus Teleostei Teleostomi Vertebrata cellular organisms
G19275EU	Fuc(a1-?)HexNAc(?1-?)Hex(?1-3)HexNAc(?1-3)[Fuc(a1-?)HexNAc(?1-?)Hex(?1-3)HexNAc(?1-4)]Gal(b1-4)Glc(?1-		Linkage defined saccharide	1770.66	Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Eukaryota Eumetazoa Euteleostomi Gnathostomata Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Teleostei Teleostomi Vertebrata cellular organisms
G76064HC	Fuc(a1-?)Fuc(a1-?)HexNAc(?1-?)GalNAc(b1-3)Gal(a1-3)Gal(b1-4)GlcNAc(b1-3)[GalNAc(b1-4)]Gal(b1-4)Glc(?1-	N-acetyllactosamine Type 2 LN a-Gal antigen	Linkage defined saccharide	1770.66	Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Eukaryota Eumetazoa Euteleostomi Gnathostomata Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Teleostei Teleostomi Vertebrata cellular organisms
G18533XI	Fuc(a1-3)GalNAc(b1-4)Gal(b1-3)Gal(b1-3)[Neu5Ac(a2-8)Neu5Ac(a2-8)Neu5Ac(a2-6)]GalNAc(a1-	O-Glycan core 1 Sialyl Tn antigen T antigen TF antigen	Linkage defined saccharide	1767.62	Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Deuterostomia Eukaryota Eumetazoa Euteleosteomorpha Euteleostomi Gnathostomata Metazoa Neopterygii Oncorhynchus mykiss Oncorhynchus Opisthokonta Osteoglossocephalai Protacanthopterygii Salmonidae Salmoniformes Salmoninae Teleostei Teleostomi Vertebrata cellular organisms
G45252ML	Fuc(a1-?)Fuc(a1-3)HexNAc(?1-?)Hex(?1-?)Gal(a1-3)Gal(b1-4)GlcNAc(b1-3)[GalNAc(b1-4)]Gal(b1-4)Glc(?1-	N-acetyllactosamine Type 2 LN a-Gal antigen	Linkage defined saccharide	1729.63	Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Eukaryota Eumetazoa Euteleostomi Gnathostomata Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Teleostei Teleostomi Vertebrata cellular organisms
G82829MF	GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex	Linkage defined saccharide	1722.65	Acanthomorphata Actinopteri Actinopterygii Adrianichthyidae Adrianichthyoidei Atherinomorphae Beloniformes Bilateria Chordata Clupeocephala Craniata Ctenosquamata Deuterostomia Euacanthomorphacea Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Gnathostomata Metazoa Neopterygii Neoteleostei Opisthokonta Oryzias melastigma Oryzias Oryziinae Osteoglossocephalai Ovalentaria Percomorphaceae Teleostei Teleostomi Vertebrata cellular organisms
G63899VB	GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan complex	Linkage defined saccharide	1722.65	Acanthomorphata Actinopteri Actinopterygii Adrianichthyidae Adrianichthyoidei Atherinomorphae Beloniformes Bilateria Chordata Clupeocephala Craniata Ctenosquamata Deuterostomia Euacanthomorphacea Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Gnathostomata Metazoa Neopterygii Neoteleostei Opisthokonta Oryzias melastigma Oryzias Oryziinae Osteoglossocephalai Ovalentaria Percomorphaceae Teleostei Teleostomi Vertebrata cellular organisms