GlyCosmos Portal

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GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	August 18, 2024

Displaying entries **151 - 175** of **1141** in total

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GlyTouCan ID ▼	IUPAC Condensed	Motifs	Subsumption Level	Monoisotopic Mass	Organisms
G87055KI	Man(a1-3)[Gal(b1-3)GlcNAc(b1-2)Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-3)]GlcNAc(?1-	N-Glycan core basic Type 1 LN	Linkage defined saccharide	1553.56	50 kb inversion clade Embryophyta Eukaryota Euphyllophyta Fabaceae Fabales Gunneridae Magnoliopsida Mesangiospermae NPAAA clade Papilionoideae Pentapetalae Phaseoleae Phaseolus vulgaris Phaseolus Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae cellular organisms eudicotyledons fabids indigoferoid/millettioid clade rosids
G87016OQ	Xyl(b1-3)Xyl(b1-4)Rha(a1-2)[Rha(a1-3)]Qui(b1-		Linkage defined saccharide	720.27	Cucurbitaceae Cucurbitales Embryophyta Eukaryota Euphyllophyta Gunneridae Luffa operculata Luffa Magnoliopsida Mesangiospermae Pentapetalae Sicyoeae Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae cellular organisms eudicotyledons fabids rosids
G86976EW	Rha(a1-3)Glc(?1-		Linkage defined saccharide	326.12	Cistanche salsa Cistanche Embryophyta Eukaryota Euphyllophyta Gunneridae Lamiales Magnoliopsida Mesangiospermae Orobanchaceae Orobancheae Pentapetalae Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae asterids cellular organisms eudicotyledons lamiids
G86895ND	Rha(b1-2)Glc(b1-2)GlcA(b1-		Linkage defined saccharide	502.15	50 kb inversion clade Embryophyta Eukaryota Euphyllophyta Fabaceae Fabales Fabeae Gunneridae Hologalegina IRL clade Lathyrus palustris subsp. pilosus Lathyrus palustris Lathyrus Magnoliopsida Mesangiospermae NPAAA clade Papilionoideae Pentapetalae Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae cellular organisms eudicotyledons fabids rosids
G86871YR	Rha(a1-2)[Glc(b1-3)]Gal		Linkage defined saccharide	488.17	Embryophyta Eukaryota Euphyllophyta Gunneridae Magnoliopsida Mesangiospermae Pentapetalae Solanaceae Solanales Solaneae Solanoideae Solanum chacoense Solanum Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae asterids cellular organisms eudicotyledons lamiids
G86688XI	Gal(?1-3)Gal(?1-?)Gal(?1-?)Ara		Linkage defined saccharide	636.21	Apiales Apiineae Araliaceae Embryophyta Eukaryota Euphyllophyta Gunneridae Hedera pastuchovii Hedera Magnoliopsida Mesangiospermae Pentapetalae Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae asterids campanulids cellular organisms eudicotyledons
G86675KY	Qui(b1-2)[Glc(b1-3)]Ara(a1-		Linkage defined saccharide	458.16	Embryophyta Eukaryota Euphyllophyta Gunneridae Hovenia dulcis Hovenia Magnoliopsida Mesangiospermae Paliureae Pentapetalae Rhamnaceae Rosales Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae cellular organisms eudicotyledons fabids rosids
G86521AY	Man(a1-3)[Man(a1-6)][Xyl(b1-2)]Man(b1-4)ManNAc-ol		Linkage defined saccharide	841.31	50 kb inversion clade Embryophyta Erythrina corallodendron Erythrina crista-galli Erythrina Eukaryota Euphyllophyta Fabaceae Fabales Gunneridae Magnoliopsida Mesangiospermae NPAAA clade Papilionoideae Pentapetalae Phaseoleae Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae cellular organisms eudicotyledons fabids indigoferoid/millettioid clade rosids
G86487TG	GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-2)Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan complex	Linkage defined saccharide	1594.59	Acalypheae Acalyphoideae Embryophyta Eukaryota Euphorbiaceae Euphyllophyta Gunneridae Magnoliopsida Malpighiales Mesangiospermae Pentapetalae Ricinus communis Ricinus Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae cellular organisms eudicotyledons fabids rosids
G86451IJ					Amniota Anacardiaceae Anacardium occidentale Anacardium Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Embryophyta Euarchontoglires Eukaryota Eumetazoa Euphyllophyta Euteleostomi Eutheria Gnathostomata Gunneridae Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Magnoliopsida Mammalia Mesangiospermae Metazoa Opisthokonta Pentapetalae Primates Sapindales Sarcopterygii Simiiformes Spermatophyta Streptophyta Streptophytina Teleostomi Tetrapoda Theria Tracheophyta Vertebrata Viridiplantae cellular organisms eudicotyledons malvids rosids
G86385RC	Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan high mannose	Linkage defined saccharide	1720.59	50 kb inversion clade Abreae Abrus precatorius Abrus Acalypheae Acalyphoideae Acalyptratae Acrogymnospermae Aesculus hippocastanum Aesculus Alpharhabdovirinae Amniota Amphiesmenoptera Amphipyrinae Anserineae Anthemideae Archelosauria Archosauria Artemisia vulgaris Artemisia Artemisiinae Arthropoda Articulavirales Artiodactyla Ascomycota Aspergillaceae Aspergillus niger Aspergillus subgen. Circumdati Aspergillus Asteraceae Asterales Asteroideae Aves BOP clade Batoidea Betula pubescens Betula Betulaceae Bilateria Bombycidae Bombycinae Bombycoidea Bombyx mori Bombyx Boreoeutheria Bos taurus Bos Bovidae Bovinae Brachycera Carpinus betulus Carpinus Caryophyllales Catarrhini Cavia porcellus Cavia Caviidae Chenopodiaceae Chenopodioideae Chondrichthyes Chordata Coelurosauria Conifers I Coturnix coturnix Coturnix Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Cyclorrhapha Deuterostomia Dicondylia Dikarya Dinosauria Dipnotetrapodomorpha Diptera Discoba Ditrysia Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Elasmobranchii Embryophyta Endopterygota Ephydroidea Eremoneura Euarchontoglires Euglenozoa Eukaryota Eumetazoa Euphorbiaceae Euphyllophyta Eurotiales Eurotiomycetes Eurotiomycetidae Euteleostomi Eutheria Fabaceae Fabales Fabeae Fagales Fungi Galliformes Galloanserae Glires Glossata Gnathostomata Gunneridae Hadeninae Haplorrhini Haploviricotina Heteroneura Hexapoda Hippocastaneae Hippocastanoideae Hologalegina Hominidae Homininae Hominoidea Homo sapiens Homo Hystricomorpha IRL clade Insecta Insthoviricetes Kinetoplastea Komagataella pastoris Komagataella Lamiales Laurasiatheria Lepidoptera Liliopsida Magnoliopsida Malpighiales Mamestra brassicae Mamestra Mammalia Mandibulata Mesangiospermae Metakinetoplastina Metazoa Monjiviricetes Mononegavirales Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha NPAAA clade Negarnaviricota Neognathae Neolepidoptera Neoptera Noctuidae Noctuoidea Obtectomera Olea europaea Olea Oleaceae Oleeae Opisthokonta Orthomyxoviridae Orthornavirae Oryza sativa Oryza Oryzeae Oryzinae Oryzoideae Panarthropoda Pancrustacea Papilionoideae Pecora Pentapetalae Perdicinae Petrosaviidae Pezizomycotina Phaffomycetaceae Phaseoleae Phaseolus vulgaris Phaseolus Phasianidae Pinaceae Pinales Pinidae Pinopsida Pinus heldreichii Pinus subgen. Pinus Pinus Pisum sativum Pisum Plusiinae Poaceae Poales Polyploviricotina Primates Protostomia Pterygota Rattus Rhabdoviridae Riboviria Ricinus communis Ricinus Rodentia Ruminantia Saccharomycetales Saccharomycetes Saccharomycotina Sapindaceae Sapindales Sarcopterygii Sauria Saurischia Sauropsida Schizophora Schizotrypanum Simiiformes Sophophora Spermatophyta Spinacia oleracea Spinacia Spodoptera frugiperda Spodoptera Streptophyta Streptophytina Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Tetronarce californica Tetronarce Theria Theropoda Torpedinidae Torpediniformes Tracheophyta Trichoplusia ni Trichoplusia Trypanosoma brucei Trypanosoma cruzi Trypanosoma Trypanosomatida Trypanosomatidae Trypanozoon Vertebrata Vesicular stomatitis virus Vesiculovirus Viridiplantae Viruses asterids campanulids cellular organisms commelinids eudicotyledons fabids indigoferoid/millettioid clade lamiids leotiomyceta malvids melanogaster group melanogaster subgroup rosids saccharomyceta unclassified Orthomyxoviridae unclassified Vesiculovirus unidentified influenza virus
G86267DW					50 kb inversion clade Arabidopsis thaliana Arabidopsis Brassicaceae Brassicales Camelineae Embryophyta Eukaryota Euphyllophyta Fabaceae Fabales Fabeae Gunneridae Hologalegina IRL clade Magnoliopsida Mesangiospermae NPAAA clade Papilionoideae Pentapetalae Phaseoleae Pisum sativum Pisum Rosales Spermatophyta Streptophyta Streptophytina Tracheophyta Ulmaceae Ulmus glabra Ulmus Vigna radiata var. radiata Vigna radiata Vigna Viridiplantae cellular organisms eudicotyledons fabids indigoferoid/millettioid clade malvids rosids
G86206ML		Mannose O-Mannose			Ascomycota Cnidaria Cyanea capillata Cyanea Cyaneidae Dikarya Embryophyta Eukaryota Eumetazoa Euphyllophyta Fungi Gunneridae Jasmineae Jasminum officinale Jasminum Lamiales Magnoliopsida Mesangiospermae Metazoa Oleaceae Opisthokonta Pentapetalae Schizosaccharomyces pombe Schizosaccharomyces Schizosaccharomycetaceae Schizosaccharomycetales Schizosaccharomycetes Scyphozoa Semaeostomeae Spermatophyta Streptophyta Streptophytina Taphrinomycotina Tracheophyta Viridiplantae asterids cellular organisms eudicotyledons lamiids
G86191DF	Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-2)Man(a1-6)]Man(a1-3)[Man(a1-2)Man(a1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan core basic	Linkage defined saccharide	2044.7	50 kb inversion clade Embryophyta Eukaryota Euphyllophyta Fabaceae Fabales Glycine max Glycine subgen. Soja Glycine Gunneridae Magnoliopsida Mesangiospermae NPAAA clade Papilionoideae Pentapetalae Phaseoleae Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae cellular organisms eudicotyledons fabids indigoferoid/millettioid clade rosids
G86156UJ	GlcNAc(b1-2)Man(a1-3)[Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-		Linkage defined saccharide	1083.4	Embryophyta Eukaryota Euphyllophyta Gunneridae Magnoliopsida Mesangiospermae Nicotiana alata Nicotiana Nicotianeae Nicotianoideae Pentapetalae Solanaceae Solanales Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae asterids cellular organisms eudicotyledons lamiids
G86053IC	Man(a1-2)Man(a1-3)[Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan high mannose	Linkage defined saccharide	1558.54	50 kb inversion clade Abreae Abrus precatorius Abrus Acalypheae Acalyphoideae Acalyptratae Acrogymnospermae Aesculus hippocastanum Aesculus Amniota Amphiesmenoptera Amphipyrinae Archelosauria Archosauria Arthropoda Articulavirales Artiodactyla Ascomycota Aspergillaceae Aspergillus niger Aspergillus subgen. Circumdati Aspergillus Aves BOP clade Betula pubescens Betula Betulaceae Bilateria Bombycidae Bombycinae Bombycoidea Bombyx mori Bombyx Boreoeutheria Bos taurus Bos Bovidae Bovinae Brachycera Brassicales Carica papaya Carica Caricaceae Carnivora Carpinus betulus Carpinus Catarrhini Chordata Coelurosauria Conifers I Coturnix coturnix Coturnix Craniata Crithidia fasciculata Crithidia Cyclorrhapha Deuterostomia Dicondylia Dikarya Dinosauria Dipnotetrapodomorpha Diptera Discoba Ditrysia Drosophila melanogaster Drosophila Drosophilidae Drosophilinae Drosophilini Ecdysozoa Embryophyta Endopterygota Ephydroidea Eremoneura Euarchontoglires Euglenozoa Eukaryota Eumetazoa Euphorbiaceae Euphyllophyta Eurotiales Eurotiomycetes Eurotiomycetidae Euteleostomi Eutheria Fabaceae Fabales Fabeae Fagales Felidae Feliformia Felinae Felis catus Felis Fungi Galliformes Galloanserae Gallus gallus Gallus Glires Glossata Gnathostomata Gunneridae Hadeninae Haplorrhini Heteroneura Hexapoda Hippocastaneae Hippocastanoideae Hologalegina Hominidae Homininae Hominoidea Homo sapiens Homo IRL clade Insecta Insthoviricetes Kinetoplastea Lamiales Laurasiatheria Leishmaniinae Lepidoptera Liliopsida Magnoliopsida Malpighiales Mamestra brassicae Mamestra Mammalia Mandibulata Mesangiospermae Metakinetoplastina Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Muscomorpha Myomorpha NPAAA clade Negarnaviricota Neognathae Neolepidoptera Neoptera Noctuidae Noctuoidea Obtectomera Olea europaea Olea Oleaceae Oleeae Opisthokonta Orthomyxoviridae Orthornavirae Oryza sativa Oryza Oryzeae Oryzinae Oryzoideae Panarthropoda Pancrustacea Papilionoideae Pecora Pentapetalae Perdicinae Petrosaviidae Pezizomycotina Phaseoleae Phaseolus vulgaris Phaseolus Phasianidae Phasianinae Pinaceae Pinales Pinidae Pinopsida Pinus heldreichii Pinus subgen. Pinus Pinus Pisum sativum Pisum Plusiinae Poaceae Poales Polyploviricotina Primates Protostomia Pterygota Rattus Riboviria Ricinus communis Ricinus Rodentia Ruminantia Sapindaceae Sapindales Sarcopterygii Sauria Saurischia Sauropsida Schizophora Simiiformes Sophophora Spermatophyta Spodoptera frugiperda Spodoptera Streptophyta Streptophytina Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Theropoda Tracheophyta Trichoplusia ni Trichoplusia Trypanosoma brucei Trypanosoma Trypanosomatida Trypanosomatidae Trypanozoon Vertebrata Viridiplantae Viruses asterids cellular organisms commelinids eudicotyledons fabids indigoferoid/millettioid clade lamiids leotiomyceta malvids melanogaster group melanogaster subgroup rosids saccharomyceta unclassified Orthomyxoviridae unclassified entries unclassified sequences unidentified influenza virus unidentified
G85877ZB	Gal(b1-4)[Fuc(a1-6)]GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)[Fuc(a1-6)]GlcNAc(b1-2)Man(a1-6)][Xyl(b1-2)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-3)]GlcNAc	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2210.81	Acer pseudoplatanus Acer Acereae Chrysophylloideae Embryophyta Ericales Eukaryota Euphyllophyta Gunneridae Hippocastanoideae Magnoliopsida Mesangiospermae Pentapetalae Platanaceae Platanus Proteales Sapindaceae Sapindales Sapotaceae Spermatophyta Streptophyta Streptophytina Synsepalum dulcificum Synsepalum Tracheophyta Viridiplantae asterids cellular organisms eudicotyledons malvids rosids
G85856RP	GlcA(b1-2)GlcA(b1-		Linkage defined saccharide	370.07	50 kb inversion clade Embryophyta Eukaryota Euphyllophyta Fabaceae Fabales Galegeae Glycyrrhiza glabra Glycyrrhiza uralensis Glycyrrhiza Gunneridae Hologalegina IRL clade Magnoliopsida Mesangiospermae NPAAA clade Papilionoideae Pentapetalae Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae cellular organisms eudicotyledons fabids rosids
G85660IO	Xyl(a1-6)Glc(b1-4)[Xyl(a1-6)]Glc(b1-4)Glc-ol		Linkage defined saccharide	770.27	50 kb inversion clade Acer pseudoplatanus Acer Acereae Amherstieae Amygdaloideae Detarioideae Embryophyta Eukaryota Euphyllophyta Fabaceae Fabales Fabeae Gunneridae Hippocastanoideae Hologalegina IRL clade Magnoliopsida Maleae Malus domestica Malus Mesangiospermae NPAAA clade Nicotiana tabacum Nicotiana Nicotianeae Nicotianoideae Papilionoideae Pentapetalae Pisum sativum Pisum Platanaceae Platanus Proteales Rosaceae Rosales Sapindaceae Sapindales Solanaceae Solanales Solaneae Solanoideae Solanum lycopersicum Solanum subgen. Lycopersicon Solanum Spermatophyta Streptophyta Streptophytina Tamarindus indica Tamarindus Tracheophyta Viridiplantae asterids cellular organisms eudicotyledons fabids lamiids malvids rosids
G85607DC	Man(a1-2)Man(a1-2)Man(a1-3)[Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(?1-		Linkage defined saccharide	1517.51	Amniota Arthropoda Articulavirales Artiodactyla Artverviricota Ascomycota Aspergillaceae Aspergillus oryzae Aspergillus subgen. Circumdati Aspergillus Avulavirinae Bilateria Blaberoidea Blattella germanica Blattella Blattellidae Blattodea Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Cricetidae Cricetinae Deuterostomia Dicondylia Dictyoptera Dikarya Dipnotetrapodomorpha Ecdysozoa Embryophyta Euarchontoglires Eukaryota Eumetazoa Euphyllophyta Eurotiales Eurotiomycetes Eurotiomycetidae Euteleostomi Eutheria Filoviridae Friend murine leukemia virus Fungi incertae sedis Fungi Gammaretrovirus Glires Gnathostomata Gunneridae Haplorrhini Haploviricotina Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Insthoviricetes Komagataella pastoris Komagataella Laurasiatheria Magnoliopsida Mammalia Mandibulata Mesangiospermae Mesocricetus auratus Mesocricetus Metazoa Monjiviricetes Mononegavirales Mucorales Mucorineae Mucoromycetes Mucoromycota Mucoromycotina Muridae Murinae Murine leukemia virus Muroidea Mus musculus Mus Mus Myomorpha Negarnaviricota Neoptera Opisthokonta Ortervirales Orthoavulavirus javaense Orthoavulavirus Orthomarburgvirus marburgense Orthomarburgvirus Orthomyxoviridae Orthoretrovirinae Orthornavirae Panarthropoda Pancrustacea Paramyxoviridae Pararnavirae Pecora Pentapetalae Pezizomycotina Phaffomycetaceae Polyneoptera Polyploviricotina Primates Protostomia Pterygota Rattus Retroviridae Revtraviricetes Rhizopodaceae Rhizopus niveus Rhizopus Riboviria Rodentia Ruminantia Saccharomycetales Saccharomycetes Saccharomycotina Sarcopterygii Simiiformes Solanaceae Solanales Solaneae Solanoideae Solanum lycopersicum Solanum subgen. Lycopersicon Solanum Spermatophyta Streptophyta Streptophytina Teleostomi Tetrapoda Theria Tracheophyta Vertebrata Viridiplantae Viruses asterids avian paramyxovirus 1 cellular organisms eudicotyledons lamiids leotiomyceta saccharomyceta unclassified Orthomyxoviridae unclassified entries unclassified sequences unidentified influenza virus unidentified
G85519TG					Ceratonia clade Ceratonia siliqua Ceratonia Embryophyta Eukaryota Euphyllophyta Fabaceae Fabales Gunneridae Magnoliopsida Mesangiospermae Pentapetalae Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae cellular organisms eudicotyledons fabids rosids
G85479PJ	GalA(a1-4)GalA(a1-4)GalA(a1-4)GalA(a1-4)GalA(?1-		Linkage defined saccharide	898.17	Aurantioideae Citrus Embryophyta Eukaryota Euphyllophyta Gunneridae Magnoliopsida Mesangiospermae Pentapetalae Rutaceae Sapindales Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae cellular organisms eudicotyledons malvids rosids
G85416AH	D-Apif(b1-2)GlcA(b1-		Linkage defined saccharide	326.08	50 kb inversion clade Embryophyta Eukaryota Euphyllophyta Fabaceae Fabales Galegeae Glycyrrhiza inflata Glycyrrhiza Gunneridae Hologalegina IRL clade Magnoliopsida Mesangiospermae NPAAA clade Papilionoideae Pentapetalae Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae cellular organisms eudicotyledons fabids rosids
G85415LL	Man(a1-6)Man(a1-6)Man(a1-	Mannose O-Mannose	Linkage defined saccharide	504.17	Actinomycetes Actinomycetota Bacteria Embryophyta Eukaryota Euphyllophyta Gunneridae Jasmineae Jasminum officinale Jasminum Lamiales Magnoliopsida Mesangiospermae Mycobacteriaceae Mycobacteriales Mycolicibacterium smegmatis Mycolicibacterium Oleaceae Pentapetalae Spermatophyta Streptophyta Streptophytina Terrabacteria group Tracheophyta Viridiplantae asterids cellular organisms eudicotyledons lamiids
G85306DM	Dig3Me(b1-4)Dig3Me(b1-		Linkage defined saccharide	306.17	Apocynaceae Asclepiadeae Asclepiadoideae Cynanchinae Cynanchum africanum Cynanchum caudatum Cynanchum Embryophyta Eukaryota Euphyllophyta Gentianales Gunneridae Magnoliopsida Mesangiospermae Pentapetalae Spermatophyta Streptophyta Streptophytina Tracheophyta Viridiplantae asterids cellular organisms eudicotyledons lamiids