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GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	August 18, 2024

Displaying entries **176 - 200** of **366** in total

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GlyTouCan ID	IUPAC Condensed ▼	Motifs	Subsumption Level	Monoisotopic Mass	Organisms
G84423SZ	GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-4)][GlcNAc(b1-2)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan complex	Linkage defined saccharide	1884.7	Amniota Archelosauria Archosauria Aves Bilateria Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Metazoa Neognathae Opisthokonta Phasianidae Phasianinae Sarcopterygii Sauria Saurischia Sauropsida Teleostomi Tetrapoda Theropoda Vertebrata cellular organisms
G20168MP	GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-4)][GlcNAc(b1-2)[GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc-ol		Linkage defined saccharide	1927.74	Amniota Archelosauria Archosauria Aves Bilateria Boreoeutheria Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Neognathae Opisthokonta Phasianidae Phasianinae Rattus Rodentia Sarcopterygii Sauria Saurischia Sauropsida Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms
G25787HL	GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-4)][GlcNAc(b1-2)[GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex	Linkage defined saccharide	1925.72	Amniota Archelosauria Archosauria Aves Bilateria Boreoeutheria Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Neognathae Opisthokonta Phasianidae Phasianinae Rattus Rodentia Sarcopterygii Sauria Saurischia Sauropsida Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms
G04717US	GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-4)][GlcNAc(b1-2)[GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan complex	Linkage defined saccharide	1925.72	Amniota Archelosauria Archosauria Aves Bilateria Boreoeutheria Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Neognathae Opisthokonta Phasianidae Phasianinae Rattus Rodentia Sarcopterygii Sauria Saurischia Sauropsida Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms
G07558KL	GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-4)][GlcNAc(b1-2)[GlcNAc(b1-4)][GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc-ol		Linkage defined saccharide	2130.82	Amniota Archelosauria Archosauria Aves Bilateria Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Metazoa Neognathae Opisthokonta Phasianidae Phasianinae Sarcopterygii Sauria Saurischia Sauropsida Teleostomi Tetrapoda Theropoda Vertebrata cellular organisms
G52187WF	GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-4)][GlcNAc(b1-2)[GlcNAc(b1-4)][GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan complex	Linkage defined saccharide	2128.8	Acanthomorphata Actinopteri Actinopterygii Amniota Archelosauria Archosauria Atherinomorphae Aves Bilateria Chordata Clupeocephala Coelurosauria Craniata Ctenosquamata Cyprinodontiformes Cyprinodontoidei Deuterostomia Dinosauria Dipnotetrapodomorpha Euacanthomorphacea Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Fundulidae Fundulus heteroclitus Fundulus Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Metazoa Neognathae Neopterygii Neoteleostei Opisthokonta Osteoglossocephalai Ovalentaria Percomorphaceae Phasianidae Phasianinae Sarcopterygii Sauria Saurischia Sauropsida Teleostei Teleostomi Tetrapoda Theropoda Vertebrata cellular organisms
G01685IB	GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-4)][GlcNAc(b1-2)[GlcNAc(b1-4)][GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex	Linkage defined saccharide	2128.8	Acanthomorphata Actinopteri Actinopterygii Amniota Archelosauria Archosauria Atherinomorphae Aves Bilateria Chordata Clupeocephala Coelurosauria Craniata Ctenosquamata Cyprinodontiformes Cyprinodontoidei Deuterostomia Dinosauria Dipnotetrapodomorpha Euacanthomorphacea Eukaryota Eumetazoa Eurypterygia Euteleosteomorpha Euteleostomi Fundulidae Fundulus heteroclitus Fundulus Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Metazoa Neognathae Neopterygii Neoteleostei Opisthokonta Osteoglossocephalai Ovalentaria Percomorphaceae Phasianidae Phasianinae Sarcopterygii Sauria Saurischia Sauropsida Teleostei Teleostomi Tetrapoda Theropoda Vertebrata cellular organisms unclassified entries unclassified sequences unidentified
G78625LN	GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-4)][GlcNAc(b1-2)[GlcNAc(b1-4)][GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan complex	Linkage defined saccharide	2128.8	Amniota Archelosauria Archosauria Aves Bilateria Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Metazoa Neognathae Opisthokonta Phasianidae Phasianinae Sarcopterygii Sauria Saurischia Sauropsida Teleostomi Tetrapoda Theropoda Vertebrata cellular organisms
G95782RB	GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-4)][GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex	Linkage defined saccharide	1925.72	Amniota Archelosauria Archosauria Aves Bilateria Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Metazoa Neognathae Opisthokonta Phasianidae Phasianinae Sarcopterygii Sauria Saurischia Sauropsida Teleostomi Tetrapoda Theropoda Vertebrata cellular organisms unclassified entries unclassified sequences unidentified
G43610AB	GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-4)][GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan complex	Linkage defined saccharide	1925.72	Amniota Archelosauria Archosauria Aves Bilateria Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Metazoa Neognathae Opisthokonta Phasianidae Phasianinae Sarcopterygii Sauria Saurischia Sauropsida Teleostomi Tetrapoda Theropoda Vertebrata cellular organisms
G77225CQ	GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-4)][GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)][GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc-ol	N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2292.87	Amniota Archelosauria Archosauria Aves Bilateria Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Metazoa Neognathae Opisthokonta Phasianidae Phasianinae Sarcopterygii Sauria Saurischia Sauropsida Teleostomi Tetrapoda Theropoda Vertebrata cellular organisms
G20508DJ	GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-4)][GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)][GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2290.86	Amniota Archelosauria Archosauria Aves Bilateria Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Metazoa Neognathae Opisthokonta Phasianidae Phasianinae Sarcopterygii Sauria Saurischia Sauropsida Teleostomi Tetrapoda Theropoda Vertebrata cellular organisms
G19894RH	GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-4)][GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc-ol		Linkage defined saccharide	1724.66	Amniota Anas platyrhynchos Anas Anatidae Anatinae Anseriformes Archelosauria Archosauria Aves Bilateria Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Metazoa Neognathae Opisthokonta Phasianidae Phasianinae Sarcopterygii Sauria Saurischia Sauropsida Teleostomi Tetrapoda Theropoda Vertebrata cellular organisms
G93500PH	GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-4)][GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan complex	Linkage defined saccharide	1722.65	Amniota Archelosauria Archosauria Aves Bilateria Boreoeutheria Catarrhini Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Neognathae Opisthokonta Phasianidae Phasianinae Primates Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms
G45256SO	GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-4)][GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex	Linkage defined saccharide	1722.65	Amniota Anas sp. Anas Anatidae Anatinae Anseriformes Archelosauria Archosauria Aves Batoidea Bilateria Boreoeutheria Catarrhini Chondrichthyes Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Elasmobranchii Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Neognathae Opisthokonta Phasianidae Phasianinae Primates Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostomi Tetrapoda Tetronarce californica Tetronarce Theria Theropoda Torpedinidae Torpediniformes Vertebrata cellular organisms unclassified Anas unclassified entries unclassified sequences unidentified
G76583MP	GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-4)][GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan complex	Linkage defined saccharide	1722.65	Amniota Archelosauria Archosauria Aves Batoidea Bilateria Boreoeutheria Catarrhini Chondrichthyes Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Elasmobranchii Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Neognathae Opisthokonta Phasianidae Phasianinae Primates Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostomi Tetrapoda Tetronarce californica Tetronarce Theria Theropoda Torpedinidae Torpediniformes Vertebrata cellular organisms
G85102OH	GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-4)][Gal(b1-4)GlcNAc(b1-2)[GlcNAc(b1-4)][GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc-ol	N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2292.87	Amniota Archelosauria Archosauria Aves Bilateria Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Metazoa Neognathae Opisthokonta Phasianidae Phasianinae Sarcopterygii Sauria Saurischia Sauropsida Teleostomi Tetrapoda Theropoda Vertebrata cellular organisms
G76397RF	GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-4)][Gal(b1-4)GlcNAc(b1-2)[GlcNAc(b1-4)][GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2290.86	Amniota Archelosauria Archosauria Aves Bilateria Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Metazoa Neognathae Opisthokonta Phasianidae Phasianinae Sarcopterygii Sauria Saurischia Sauropsida Teleostomi Tetrapoda Theropoda Vertebrata cellular organisms
G20030CU	GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-2)[GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan complex	Linkage defined saccharide	1722.65	Amniota Archelosauria Archosauria Aves Bilateria Boreoeutheria Catarrhini Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Neognathae Opisthokonta Phasianidae Phasianinae Primates Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms
G78509IY	GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-2)[GlcNAc(b1-4)][GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex	Linkage defined saccharide	1925.72	Amniota Archelosauria Archosauria Aves Bilateria Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Metazoa Neognathae Opisthokonta Phasianidae Phasianinae Sarcopterygii Sauria Saurischia Sauropsida Teleostomi Tetrapoda Theropoda Vertebrata cellular organisms
G46415RP	GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-2)[GlcNAc(b1-4)][GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan complex	Linkage defined saccharide	1925.72	Amniota Archelosauria Archosauria Aves Bilateria Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Metazoa Neognathae Opisthokonta Phasianidae Phasianinae Sarcopterygii Sauria Saurischia Sauropsida Teleostomi Tetrapoda Theropoda Vertebrata cellular organisms
G16717IV	GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc-ol		Linkage defined saccharide	1521.58	Amniota Anas platyrhynchos Anas Anatidae Anatinae Anseriformes Archelosauria Archosauria Articulavirales Artiodactyla Aves Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Insthoviricetes Laurasiatheria Mammalia Metazoa Negarnaviricota Neognathae Opisthokonta Orthomyxoviridae Orthornavirae Pecora Phasianidae Phasianinae Polyploviricotina Primates Riboviria Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostomi Tetrapoda Theria Theropoda Vertebrata Viruses cellular organisms unclassified Orthomyxoviridae unidentified influenza virus
G29011JC	GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan complex	Linkage defined saccharide	1519.57	Amniota Archelosauria Archosauria Artiodactyla Aves Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Neognathae Opisthokonta Pecora Phasianidae Phasianinae Primates Rattus norvegicus Rattus Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms
G75584LU	GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex	Linkage defined saccharide	1519.57	Amniota Anas sp. Anas Anatidae Anatinae Anseriformes Archelosauria Archosauria Articulavirales Artiodactyla Aves Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Insthoviricetes Laurasiatheria Mammalia Metazoa Negarnaviricota Neognathae Opisthokonta Orthomyxoviridae Orthornavirae Pecora Phasianidae Phasianinae Polyploviricotina Primates Riboviria Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostomi Tetrapoda Theria Theropoda Vertebrata Viruses cellular organisms unclassified Anas unclassified Orthomyxoviridae unidentified influenza virus
G65415JG	GlcNAc(b1-2)[GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan complex	Linkage defined saccharide	1519.57	Amniota Archelosauria Archosauria Artiodactyla Aves Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Laurasiatheria Mammalia Metazoa Neognathae Opisthokonta Pecora Phasianidae Phasianinae Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms