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Guidelines

MIRAGE

autophagy related 4B, cysteine peptidase

Summary

Gene Symbol

Atg4b

Organism

Mus musculus (house mouse)

External Links

NCBI Gene

66615

PubChem

66615

Alliance of Genome Resources

MGI:1913865

Annotation

Keyword

Acetylation
Autophagy
Cytoplasm
Cytoplasmic vesicle
Disulfide bond
Endoplasmic reticulum
Glycoprotein
Isopeptide bond
Mitochondrion
Phosphoprotein
Protein transport
Proteomics identification
Reference proteome
S-nitrosylation
Thiol protease
Ubl conjugation
Ubl conjugation pathway

Proteins

Displaying **all 2** entries
UniProt	Protein Name
Q8BGE6	AUT-like 1 cysteine endopeptidase Autophagy-related cysteine endopeptidase 1 Autophagy-related protein 4 homolog B
A0A0R4J065

Annotation information from UniProt.

Gene Ontology (GO)

Displaying entries **1 - 5** of 23 in total

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GO Term	Evidence Code	PMID
autophagosome assembly	ISO
autophagosome assembly	IMP	25578879
autophagosome assembly	IBA
mitophagy	ISS
mitophagy	ISO

GO Hierarchy

biological process

metabolic process [inference]

nitrogen compound metabolic process [inference]

organonitrogen compound metabolic process [inference]

protein metabolic process [inference]

protein processing

protein modification process [inference]

protein delipidation

protein maturation [inference]

protein processing

catabolic process [inference]

cellular catabolic process [inference]

autophagy of mitochondrion [inference]

mitophagy

macroautophagy [inference]

nucleophagy [inference]

piecemeal microautophagy of the nucleus

piecemeal microautophagy of the nucleus

cellular metabolic process [inference]

cellular catabolic process [inference]

autophagy of mitochondrion [inference]

mitophagy

macroautophagy [inference]

nucleophagy [inference]

piecemeal microautophagy of the nucleus

piecemeal microautophagy of the nucleus

primary metabolic process [inference]

protein metabolic process [inference]

protein processing

protein modification process [inference]

protein delipidation

protein maturation [inference]

protein processing

organic substance metabolic process [inference]

macromolecule metabolic process [inference]

protein metabolic process [inference]

protein processing

protein modification process [inference]

protein delipidation

protein maturation [inference]

protein processing

macromolecule modification [inference]

protein modification process [inference]

protein delipidation

organonitrogen compound metabolic process [inference]

protein metabolic process [inference]

protein processing

protein modification process [inference]

protein delipidation

protein maturation [inference]

protein processing

cellular process [inference]

cell communication [inference]

cellular response to extracellular stimulus [inference]

cellular response to nutrient levels [inference]

cellular response to starvation

cellular metabolic process [inference]

cellular catabolic process [inference]

autophagy of mitochondrion [inference]

mitophagy

macroautophagy [inference]

nucleophagy [inference]

piecemeal microautophagy of the nucleus

piecemeal microautophagy of the nucleus

cellular localization [inference]

cellular macromolecule localization [inference]

protein localization [inference]

protein localization to phagophore assembly site

establishment of protein localization [inference]

protein transport

cellular response to stimulus [inference]

cellular response to extracellular stimulus [inference]

cellular response to nutrient levels [inference]

cellular response to starvation

cellular response to stress [inference]

cellular response to starvation

process utilizing autophagic mechanism [inference]

autophagy of mitochondrion [inference]

mitophagy

macroautophagy [inference]

nucleophagy [inference]

piecemeal microautophagy of the nucleus

piecemeal microautophagy of the nucleus

cellular component organization or biogenesis [inference]

cellular component organization [inference]

organelle organization [inference]

mitochondrion organization [inference]

autophagy of mitochondrion [inference]

mitophagy

vacuole organization [inference]

autophagosome organization [inference]

autophagosome assembly

organelle assembly [inference]

autophagosome assembly

organelle disassembly [inference]

autophagy of mitochondrion [inference]

mitophagy

cellular component disassembly [inference]

organelle disassembly [inference]

autophagy of mitochondrion [inference]

mitophagy

cellular component assembly [inference]

organelle assembly [inference]

autophagosome assembly

developmental process [inference]

anatomical structure development [inference]

tissue development [inference]

biomineral tissue development [inference]

otolith mineralization [inference]

otolith mineralization completed early in development

response to stimulus [inference]

response to stress [inference]

cellular response to stress [inference]

cellular response to starvation

response to starvation [inference]

cellular response to starvation

response to external stimulus [inference]

response to extracellular stimulus [inference]

response to nutrient levels [inference]

cellular response to nutrient levels [inference]

cellular response to starvation

response to starvation [inference]

cellular response to starvation

cellular response to extracellular stimulus [inference]

cellular response to nutrient levels [inference]

cellular response to starvation

cellular response to external stimulus [inference]

cellular response to extracellular stimulus [inference]

cellular response to nutrient levels [inference]

cellular response to starvation

cellular response to stimulus [inference]

cellular response to extracellular stimulus [inference]

cellular response to nutrient levels [inference]

cellular response to starvation

cellular response to stress [inference]

cellular response to starvation

localization [inference]

macromolecule localization [inference]

cellular macromolecule localization [inference]

protein localization [inference]

protein localization to phagophore assembly site

establishment of protein localization [inference]

protein transport

establishment of localization [inference]

transport [inference]

organic substance transport [inference]

protein transport

nitrogen compound transport [inference]

protein transport

establishment of protein localization [inference]

protein transport

cellular localization [inference]

cellular macromolecule localization [inference]

protein localization [inference]

protein localization to phagophore assembly site

establishment of protein localization [inference]

protein transport

biological regulation [inference]

regulation of biological process [inference]

regulation of metabolic process [inference]

positive regulation of metabolic process [inference]

positive regulation of catabolic process [inference]

positive regulation of protein catabolic process

positive regulation of macromolecule metabolic process [inference]

positive regulation of protein metabolic process [inference]

positive regulation of protein catabolic process

positive regulation of nitrogen compound metabolic process [inference]

positive regulation of protein metabolic process [inference]

positive regulation of protein catabolic process

regulation of catabolic process [inference]

positive regulation of catabolic process [inference]

positive regulation of protein catabolic process

regulation of protein catabolic process [inference]

positive regulation of protein catabolic process

regulation of nitrogen compound metabolic process [inference]

positive regulation of nitrogen compound metabolic process [inference]

positive regulation of protein metabolic process [inference]

positive regulation of protein catabolic process

regulation of protein metabolic process [inference]

regulation of protein catabolic process [inference]

positive regulation of protein catabolic process

positive regulation of protein metabolic process [inference]

positive regulation of protein catabolic process

regulation of macromolecule metabolic process [inference]

positive regulation of macromolecule metabolic process [inference]

positive regulation of protein metabolic process [inference]

positive regulation of protein catabolic process

regulation of protein metabolic process [inference]

regulation of protein catabolic process [inference]

positive regulation of protein catabolic process

positive regulation of protein metabolic process [inference]

positive regulation of protein catabolic process

regulation of primary metabolic process [inference]

regulation of protein metabolic process [inference]

regulation of protein catabolic process [inference]

positive regulation of protein catabolic process

positive regulation of protein metabolic process [inference]

positive regulation of protein catabolic process

positive regulation of biological process [inference]

positive regulation of metabolic process [inference]

positive regulation of catabolic process [inference]

positive regulation of protein catabolic process

positive regulation of macromolecule metabolic process [inference]

positive regulation of protein metabolic process [inference]

positive regulation of protein catabolic process

positive regulation of nitrogen compound metabolic process [inference]

positive regulation of protein metabolic process [inference]

positive regulation of protein catabolic process

Displaying entries **1 - 5** of 8 in total

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GO Term	Evidence Code	PMID
autophagosome membrane	ISO
cytoplasm	IDA	14530254
cytoplasm	IBA
mitochondrion	IEA
autophagosome	IEA

GO Hierarchy

cellular component

cellular anatomical entity [inference]

membrane [inference]

organelle membrane [inference]

bounding membrane of organelle [inference]

vacuolar membrane [inference]

autophagosome membrane

organelle [inference]

membrane-bounded organelle [inference]

vesicle [inference]

intracellular vesicle [inference]

cytoplasmic vesicle

intracellular membrane-bounded organelle [inference]

mitochondrion

vacuole [inference]

autophagosome

endoplasmic reticulum

intracellular vesicle [inference]

cytoplasmic vesicle

intracellular organelle [inference]

intracellular membrane-bounded organelle [inference]

mitochondrion

vacuole [inference]

autophagosome

endoplasmic reticulum

intracellular vesicle [inference]

cytoplasmic vesicle

Displaying entries **1 - 5** of 12 in total

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GO Term	Evidence Code	PMID
endopeptidase activity	ISO
cysteine-type endopeptidase activity	ISO
cysteine-type endopeptidase activity	IBA
protein binding	IPI	14530254 21551061 23093945
peptidase activity	IDA	14530254

GO Hierarchy

molecular function

catalytic activity [inference]

hydrolase activity [inference]

peptidase activity

endopeptidase activity

cysteine-type endopeptidase activity

cysteine-type peptidase activity

cysteine-type endopeptidase activity

protein-phosphatidylethanolamide deconjugating activity

catalytic activity, acting on a protein [inference]

peptidase activity

endopeptidase activity

cysteine-type endopeptidase activity

cysteine-type peptidase activity

cysteine-type endopeptidase activity

protein-phosphatidylethanolamide deconjugating activity

binding [inference]

protein binding

scaffold protein binding

Gene Ontology information from NCBI Gene and UniProt.

OrthoDB (Group)

Group level

Eukaryota

Group Name

cysteine protease

Functional Category

E: Amino acid transport and metabolism
O: Posttranslational modification, protein turnover, chaperones
T: Signal transduction mechanisms

Displaying **all 3** entries
Gene Ontology
hydrolase activity
protein-phosphatidylethanolamide deconjugating activity
proteolysis

Displaying **all 3** entries
InterPro
Papain-like cysteine peptidase superfamily
Peptidase C54, catalytic domain
Peptidase C54

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International Collaboration

GlyCosmos is a member of the GlySpace Alliance together with GlyGen and Glycomics@ExPASy.

Acknowledgements

Supported by JST NBDC Grant Number JPMJND2204

Partly supported by NIH Common Fund Grant #1U01GM125267-01

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Last updated: April 6, 2026