GlyCosmos Portal

Annotation

Keyword

Disulfide bond
Glycoprotein
Glycosyltransferase
Golgi apparatus
Lipid metabolism
Manganese
Metal-binding
Proteomics identification
Reference proteome
Secreted
Signal
Signal-anchor
Transmembrane helix

Proteins

Displaying **all 4** entries
UniProt	Protein Name
O60513	Beta-N-acetylglucosaminyl-glycolipid beta-1,4-galactosyltransferase Lactotriaosylceramide beta-1,4-galactosyltransferase N-acetyllactosamine synthase Nal synthase UDP-Gal:beta-GlcNAc beta-1,4-galactosyltransferase 4 UDP-galactose:beta-N-acetylglucosamine beta-1,4-galactosyltransferase 4
B3KM35
E7ETS9
B2RAZ5

Annotation information from UniProt.

Gene Ontology (GO)

Displaying entries **1 - 5** of 7 in total

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GO Term	Evidence Code	PMID
lactosylceramide biosynthetic process	IDA	9792633
carbohydrate metabolic process	IEA
protein glycosylation	IEA
membrane lipid metabolic process	TAS	9792633
keratan sulfate biosynthetic process	IDA	17690104

GO Hierarchy

biological process

metabolic process [inference]

nitrogen compound metabolic process [inference]

cellular nitrogen compound metabolic process [inference]

cellular nitrogen compound biosynthetic process [inference]

amide biosynthetic process [inference]

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

amide metabolic process [inference]

ceramide metabolic process [inference]

lactosylceramide metabolic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

amide biosynthetic process [inference]

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

organonitrogen compound metabolic process [inference]

aminoglycan metabolic process [inference]

aminoglycan biosynthetic process [inference]

glycosaminoglycan biosynthetic process [inference]

keratan sulfate biosynthetic process

glycosaminoglycan metabolic process [inference]

glycosaminoglycan biosynthetic process [inference]

keratan sulfate biosynthetic process

mucopolysaccharide metabolic process [inference]

keratan sulfate metabolic process [inference]

keratan sulfate biosynthetic process

sphingolipid metabolic process [inference]

ceramide metabolic process [inference]

lactosylceramide metabolic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

glycosphingolipid metabolic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

lactosylceramide metabolic process [inference]

lactosylceramide biosynthetic process

sphingolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

protein metabolic process [inference]

protein modification process [inference]

protein glycosylation

organonitrogen compound biosynthetic process [inference]

aminoglycan biosynthetic process [inference]

glycosaminoglycan biosynthetic process [inference]

keratan sulfate biosynthetic process

sphingolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

biosynthetic process [inference]

cellular biosynthetic process [inference]

macromolecule biosynthetic process [inference]

aminoglycan biosynthetic process [inference]

glycosaminoglycan biosynthetic process [inference]

keratan sulfate biosynthetic process

cellular nitrogen compound biosynthetic process [inference]

amide biosynthetic process [inference]

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

sulfur compound biosynthetic process [inference]

keratan sulfate biosynthetic process

membrane lipid biosynthetic process [inference]

glycolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

sphingolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

organic substance biosynthetic process [inference]

lipid biosynthetic process [inference]

membrane lipid biosynthetic process [inference]

glycolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

sphingolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

macromolecule biosynthetic process [inference]

aminoglycan biosynthetic process [inference]

glycosaminoglycan biosynthetic process [inference]

keratan sulfate biosynthetic process

carbohydrate derivative biosynthetic process [inference]

aminoglycan biosynthetic process [inference]

glycosaminoglycan biosynthetic process [inference]

keratan sulfate biosynthetic process

glycolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

organonitrogen compound biosynthetic process [inference]

aminoglycan biosynthetic process [inference]

glycosaminoglycan biosynthetic process [inference]

keratan sulfate biosynthetic process

sphingolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

cellular metabolic process [inference]

sulfur compound metabolic process [inference]

keratan sulfate metabolic process [inference]

keratan sulfate biosynthetic process

sulfur compound biosynthetic process [inference]

keratan sulfate biosynthetic process

cellular nitrogen compound metabolic process [inference]

cellular nitrogen compound biosynthetic process [inference]

amide biosynthetic process [inference]

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

cellular biosynthetic process [inference]

macromolecule biosynthetic process [inference]

aminoglycan biosynthetic process [inference]

glycosaminoglycan biosynthetic process [inference]

keratan sulfate biosynthetic process

cellular nitrogen compound biosynthetic process [inference]

amide biosynthetic process [inference]

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

sulfur compound biosynthetic process [inference]

keratan sulfate biosynthetic process

membrane lipid biosynthetic process [inference]

glycolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

sphingolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

cellular lipid metabolic process [inference]

membrane lipid metabolic process

glycolipid metabolic process [inference]

glycosphingolipid metabolic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

lactosylceramide metabolic process [inference]

lactosylceramide biosynthetic process

glycolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

sphingolipid metabolic process [inference]

ceramide metabolic process [inference]

lactosylceramide metabolic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

glycosphingolipid metabolic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

lactosylceramide metabolic process [inference]

lactosylceramide biosynthetic process

sphingolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

membrane lipid biosynthetic process [inference]

glycolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

sphingolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

primary metabolic process [inference]

carbohydrate metabolic process

lipid metabolic process [inference]

lipid biosynthetic process [inference]

membrane lipid biosynthetic process [inference]

glycolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

sphingolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

cellular lipid metabolic process [inference]

membrane lipid metabolic process

glycolipid metabolic process [inference]

glycosphingolipid metabolic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

lactosylceramide metabolic process [inference]

lactosylceramide biosynthetic process

glycolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

sphingolipid metabolic process [inference]

ceramide metabolic process [inference]

lactosylceramide metabolic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

glycosphingolipid metabolic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

lactosylceramide metabolic process [inference]

lactosylceramide biosynthetic process

sphingolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

membrane lipid biosynthetic process [inference]

glycolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

sphingolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

liposaccharide metabolic process [inference]

glycolipid metabolic process [inference]

glycosphingolipid metabolic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

lactosylceramide metabolic process [inference]

lactosylceramide biosynthetic process

glycolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

protein metabolic process [inference]

protein modification process [inference]

protein glycosylation

glycosylation

macromolecule glycosylation [inference]

protein glycosylation

organic substance metabolic process [inference]

carbohydrate metabolic process

lipid metabolic process [inference]

lipid biosynthetic process [inference]

membrane lipid biosynthetic process [inference]

glycolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

sphingolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

cellular lipid metabolic process [inference]

membrane lipid metabolic process

glycolipid metabolic process [inference]

glycosphingolipid metabolic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

lactosylceramide metabolic process [inference]

lactosylceramide biosynthetic process

glycolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

sphingolipid metabolic process [inference]

ceramide metabolic process [inference]

lactosylceramide metabolic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

glycosphingolipid metabolic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

lactosylceramide metabolic process [inference]

lactosylceramide biosynthetic process

sphingolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

membrane lipid biosynthetic process [inference]

glycolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

sphingolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

liposaccharide metabolic process [inference]

glycolipid metabolic process [inference]

glycosphingolipid metabolic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

lactosylceramide metabolic process [inference]

lactosylceramide biosynthetic process

glycolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

macromolecule metabolic process [inference]

aminoglycan metabolic process [inference]

aminoglycan biosynthetic process [inference]

glycosaminoglycan biosynthetic process [inference]

keratan sulfate biosynthetic process

glycosaminoglycan metabolic process [inference]

glycosaminoglycan biosynthetic process [inference]

keratan sulfate biosynthetic process

mucopolysaccharide metabolic process [inference]

keratan sulfate metabolic process [inference]

keratan sulfate biosynthetic process

macromolecule biosynthetic process [inference]

aminoglycan biosynthetic process [inference]

glycosaminoglycan biosynthetic process [inference]

keratan sulfate biosynthetic process

protein metabolic process [inference]

protein modification process [inference]

protein glycosylation

macromolecule modification [inference]

protein modification process [inference]

protein glycosylation

macromolecule glycosylation [inference]

protein glycosylation

carbohydrate derivative metabolic process [inference]

aminoglycan metabolic process [inference]

aminoglycan biosynthetic process [inference]

glycosaminoglycan biosynthetic process [inference]

keratan sulfate biosynthetic process

glycosaminoglycan metabolic process [inference]

glycosaminoglycan biosynthetic process [inference]

keratan sulfate biosynthetic process

mucopolysaccharide metabolic process [inference]

keratan sulfate metabolic process [inference]

keratan sulfate biosynthetic process

carbohydrate derivative biosynthetic process [inference]

aminoglycan biosynthetic process [inference]

glycosaminoglycan biosynthetic process [inference]

keratan sulfate biosynthetic process

glycolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

liposaccharide metabolic process [inference]

glycolipid metabolic process [inference]

glycosphingolipid metabolic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

lactosylceramide metabolic process [inference]

lactosylceramide biosynthetic process

glycolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

organonitrogen compound metabolic process [inference]

aminoglycan metabolic process [inference]

aminoglycan biosynthetic process [inference]

glycosaminoglycan biosynthetic process [inference]

keratan sulfate biosynthetic process

glycosaminoglycan metabolic process [inference]

glycosaminoglycan biosynthetic process [inference]

keratan sulfate biosynthetic process

mucopolysaccharide metabolic process [inference]

keratan sulfate metabolic process [inference]

keratan sulfate biosynthetic process

sphingolipid metabolic process [inference]

ceramide metabolic process [inference]

lactosylceramide metabolic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

glycosphingolipid metabolic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

lactosylceramide metabolic process [inference]

lactosylceramide biosynthetic process

sphingolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

protein metabolic process [inference]

protein modification process [inference]

protein glycosylation

organonitrogen compound biosynthetic process [inference]

aminoglycan biosynthetic process [inference]

glycosaminoglycan biosynthetic process [inference]

keratan sulfate biosynthetic process

sphingolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

organic substance biosynthetic process [inference]

lipid biosynthetic process [inference]

membrane lipid biosynthetic process [inference]

glycolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

sphingolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

macromolecule biosynthetic process [inference]

aminoglycan biosynthetic process [inference]

glycosaminoglycan biosynthetic process [inference]

keratan sulfate biosynthetic process

carbohydrate derivative biosynthetic process [inference]

aminoglycan biosynthetic process [inference]

glycosaminoglycan biosynthetic process [inference]

keratan sulfate biosynthetic process

glycolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

organonitrogen compound biosynthetic process [inference]

aminoglycan biosynthetic process [inference]

glycosaminoglycan biosynthetic process [inference]

keratan sulfate biosynthetic process

sphingolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

cellular process [inference]

cellular metabolic process [inference]

sulfur compound metabolic process [inference]

keratan sulfate metabolic process [inference]

keratan sulfate biosynthetic process

sulfur compound biosynthetic process [inference]

keratan sulfate biosynthetic process

cellular nitrogen compound metabolic process [inference]

cellular nitrogen compound biosynthetic process [inference]

amide biosynthetic process [inference]

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

cellular biosynthetic process [inference]

macromolecule biosynthetic process [inference]

aminoglycan biosynthetic process [inference]

glycosaminoglycan biosynthetic process [inference]

keratan sulfate biosynthetic process

cellular nitrogen compound biosynthetic process [inference]

amide biosynthetic process [inference]

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

sulfur compound biosynthetic process [inference]

keratan sulfate biosynthetic process

membrane lipid biosynthetic process [inference]

glycolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

sphingolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

cellular lipid metabolic process [inference]

membrane lipid metabolic process

glycolipid metabolic process [inference]

glycosphingolipid metabolic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

lactosylceramide metabolic process [inference]

lactosylceramide biosynthetic process

glycolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

sphingolipid metabolic process [inference]

ceramide metabolic process [inference]

lactosylceramide metabolic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

glycosphingolipid metabolic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

lactosylceramide metabolic process [inference]

lactosylceramide biosynthetic process

sphingolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

membrane lipid biosynthetic process [inference]

glycolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

sphingolipid biosynthetic process [inference]

glycosphingolipid biosynthetic process [inference]

lactosylceramide biosynthetic process

ceramide biosynthetic process [inference]

lactosylceramide biosynthetic process

Displaying entries **1 - 5** of 7 in total

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GO Term	Evidence Code	PMID
Golgi membrane	TAS
Golgi membrane	IDA	32827291
extracellular region	IEA
Golgi apparatus	IDA
Golgi apparatus	IBA

GO Hierarchy

cellular component

cellular anatomical entity [inference]

extracellular region

membrane

organelle membrane [inference]

Golgi cisterna membrane

bounding membrane of organelle [inference]

Golgi membrane

organelle [inference]

membrane-bounded organelle [inference]

intracellular membrane-bounded organelle [inference]

Golgi apparatus

intracellular organelle [inference]

intracellular membrane-bounded organelle [inference]

Golgi apparatus

Displaying entries **1 - 5** of 6 in total

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GO Term	Evidence Code	PMID
N-acetyllactosamine synthase activity	IDA	9792633
protein binding	IPI	32827291
galactosyltransferase activity	IBA
glycosyltransferase activity
UDP-galactosyltransferase activity	IDA	12511560 17690104 32827291 9792633

GO Hierarchy

molecular function

catalytic activity [inference]

transferase activity [inference]

glycosyltransferase activity

UDP-glycosyltransferase activity [inference]

UDP-galactosyltransferase activity

N-acetyllactosamine synthase activity

hexosyltransferase activity [inference]

galactosyltransferase activity

UDP-galactosyltransferase activity

N-acetyllactosamine synthase activity

binding [inference]

protein binding

ion binding [inference]

cation binding [inference]

metal ion binding

Gene Ontology information from NCBI Gene and UniProt.

GlycoGene Database (GGDB)

GGDB ID

gg080

Gene Symbol

B4GALT4

Reactions

Displaying **all 3** entries
Donor	Acceptor	Reducing terminal(Acceptor)	Product	Reducing terminal(Product)	Reference
UDP-Gal	G55302WQ	R	G08863NS	R	17529980
UDP-Gal	G06239NA	Cer	G58896AZ	Cer	9792633
UDP-Gal	G21828TB	Cer	G24366HE	Cer	9792633

Displaying entries **16 - 20** of 20 in total

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Donor	Acceptor	Reducing terminal(Acceptor)	Product	Reducing terminal(Product)	Reference
UDP-Gal	G00033MO	pNP	G64973KT	pNP	9857011
UDP-Gal	G21828TB	Cer	G24366HE	Cer	9792633
UDP-Gal	G82440PA	octyl	G00887EN	octyl	9857011
UDP-Gal	G85856KC		G28052FT		9792633
UDP-Gal	G43211QV		G71893QN		12511560

Displaying entries **1 - 5** of 10 in total

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Donor	Acceptor	Reducing terminal(Acceptor)	Reference
UDP-Gal	G68544GH	[alpha]-1-Methyl	9792633
UDP-Gal	G65889KE	[beta]-1-4-methyl-umbelliferyl	9792633
UDP-Gal	G61418DU	[beta]-1-4-methyl-umbelliferyl	9792633
UDP-Gal	G49108TO		9792633
UDP-Gal	G65889KE	[beta]-1-o-Nitrophenyl	9792633