GlyCosmos Portal

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GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	August 18, 2024

Displaying entries **1451 - 1475** of **1810** in total

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GlyTouCan ID	IUPAC Condensed	Motifs	Subsumption Level	Monoisotopic Mass ▼	Organisms
G59508TM	Gal(b1-3)GalNAc(b1-4)Gal3S(b1-4)Glc(b1-	Glucosylceramide	Linkage defined saccharide	787.21	Amniota Artiodactyla Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Primates Rattus Rodentia Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G36161EP			Linkage defined saccharide	774.22	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G42318RN	Neu5Ac(a2-6)Gal(b1-4)GlcNAc6P		Linkage defined saccharide	754.2	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G17401LE	Glc(b1-4)GlcNAc(b1-4)Gal(b1-3)GalNAc-ol		Linkage defined saccharide	750.29	Amniota Artiodactyla Bilateria Boreoeutheria Bovidae Capra hircus Capra Caprinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G13947MP	Man(a1-6)Man(b1-4)GlcNAc(b1-4)GlcNAc-ol		Linkage defined saccharide	750.29	Aculeata Alveolata Amniota Anas platyrhynchos Anas Anatidae Anatinae Anseriformes Anthophila Apidae Apinae Apini Apis mellifera Apis Apocrita Apoidea Archelosauria Archosauria Arthropoda Artiodactyla Aves Bilateria Boreoeutheria Catarrhini Chordata Ciliophora Coelurosauria Craniata Deuterostomia Dicondylia Dinosauria Dipnotetrapodomorpha Ecdysozoa Endopterygota Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Glires Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Hymenoptera Hymenostomatida Insecta Intramacronucleata Laurasiatheria Mammalia Mandibulata Metazoa Muridae Murinae Muroidea Myomorpha Neognathae Neoptera Oligohymenophorea Opisthokonta Panarthropoda Pancrustacea Primates Protostomia Pterygota Rattus Rodentia Sar Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrahymena pyriformis Tetrahymena Tetrahymenidae Tetrahymenina Tetrapoda Theria Theropoda Vertebrata cellular organisms
G18132YR	Gal(b1-3)[Gal(b1-4)GlcNAc(b1-6)]GalNAc-ol	N-acetyllactosamine Type 2 LN	Linkage defined saccharide	750.29	Amniota Apodidae Apodiformes Apodinae Archelosauria Archosauria Artiodactyla Aves Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Caprimulgimorphae Caprinae Catarrhini Chordata Coelurosauria Collocalia Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Neognathae Opisthokonta Ovis aries Ovis Pecora Phasianidae Phasianinae Primates Rattus Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms
G54461SS	Gal(b1-4)GlcNAc(b1-3)Gal(b1-3)GalNAc-ol	N-acetyllactosamine Type 2 LN	Linkage defined saccharide	750.29	Amniota Artiodactyla Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Digenea Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Lophotrochozoa Mammalia Metazoa Opisthokonta Platyhelminthes Primates Protostomia Sarcopterygii Schistosoma mansoni Schistosoma Schistosomatidae Schistosomatoidea Simiiformes Spiralia Strigeidida Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Trematoda Vertebrata cellular organisms
G64973KT	Gal(b1-3)[Gal(b1-4)GlcNAc(b1-6)]GalNAc(a1-	N-acetyllactosamine O-Glycan core 2 Type 2 LN	Linkage defined saccharide	748.27	Actinopteri Actinopterygii Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Caprinae Catarrhini Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Ovis aries Ovis Pecora Primates Rattus norvegicus Rattus Rodentia Ruminantia Sarcopterygii Simiiformes Teleostei Teleostomi Tetrapoda Theria Vertebrata artificial sequences cellular organisms other entries other sequences synthetic construct
G17075EV	Man(a1-6)Man(b1-4)GlcNAc(b1-4)GlcNAc		Linkage defined saccharide	748.27	Aculeata Amniota Amphiesmenoptera Amphipyrinae Anthophila Apidae Apinae Apini Apis mellifera Apis Apocrita Apoidea Arthropoda Artiodactyla Ascomycota Bilateria Boreoeutheria Bovidae Capra hircus Capra Caprinae Carnivora Catarrhini Chordata Craniata Deuterostomia Dicondylia Dikarya Dipnotetrapodomorpha Ditrysia Ecdysozoa Endopterygota Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Felidae Feliformia Felinae Felis catus Felis Fungi Glires Glossata Gnathostomata Haplorrhini Heteroneura Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Hymenoptera Insecta Laurasiatheria Lepidoptera Mammalia Mandibulata Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neolepidoptera Neoptera Noctuidae Noctuoidea Obtectomera Opisthokonta Ovis aries Ovis Panarthropoda Pancrustacea Pecora Primates Protostomia Pterygota Rodentia Ruminantia Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales Saccharomycetes Saccharomycotina Sarcopterygii Simiiformes Spodoptera frugiperda Spodoptera Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms saccharomyceta
G40161ME	Man(a1-3)Man(b1-4)GlcNAc(b1-4)GlcNAc		Linkage defined saccharide	748.27	50 kb inversion clade Amniota Artiodactyla Ascomycota Bilateria Boreoeutheria Carnivora Chordata Craniata Deuterostomia Dikarya Dipnotetrapodomorpha Embryophyta Euarchontoglires Eukaryota Eumetazoa Euphyllophyta Euteleostomi Eutheria Fabaceae Fabales Felidae Feliformia Felinae Felis catus Felis Fungi Glires Gnathostomata Gunneridae Laurasiatheria Magnoliopsida Mammalia Mesangiospermae Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha NPAAA clade Opisthokonta Papilionoideae Pentapetalae Phaseoleae Rodentia Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales Saccharomycetes Saccharomycotina Sarcopterygii Spermatophyta Streptophyta Streptophytina Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Tracheophyta Vertebrata Vigna radiata var. radiata Vigna radiata Vigna Viridiplantae cellular organisms eudicotyledons fabids indigoferoid/millettioid clade rosids saccharomyceta
G41884AT	GlcNAc(b1-3)[Gal(b1-4)GlcNAc(b1-6)]Gal	N-acetyllactosamine Type 2 LN	Linkage defined saccharide	748.27	Amniota Artiodactyla Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Sarcopterygii Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G05108UT	Man(a1-6)Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-		Linkage defined saccharide	748.27	Aculeata Agelenidae Agelenopsis aperta Agelenopsis Alveolata Amniota Amphiesmenoptera Amphipyrinae Ananas comosus Ananas Anas platyrhynchos Anas sp. Anas Anatidae Anatinae Anseriformes Anthophila Apidae Apinae Apini Apis mellifera Apis Apocrita Apoidea Arachnida Araneae Araneomorphae Archelosauria Archosauria Armoracia rusticana Armoracia Arthropoda Artiodactyla Ascomycota Aves Bilateria Bombycidae Bombycinae Bombycoidea Bombyx mori Bombyx Boreoeutheria Bos taurus Bos Bovidae Bovinae Brassicaceae Brassicales Bromeliaceae Bromelioideae Capra hircus Capra Caprinae Cardamineae Carica papaya Carica Caricaceae Carnivora Catarrhini Chelicerata Chordata Ciliophora Coelurosauria Coturnix coturnix Coturnix Craniata Deuterostomia Dicondylia Dikarya Dinosauria Dipnotetrapodomorpha Ditrysia Ecdysozoa Embryophyta Endopterygota Entelegynae Euarchontoglires Eukaryota Eumetazoa Euphyllophyta Euteleostomi Eutheria Felidae Feliformia Felinae Felis catus Felis Fungi Galliformes Galloanserae Gallus gallus Gallus Glires Glossata Gnathostomata Gunneridae Hadeninae Haplorrhini Heteroneura Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Hymenoptera Hymenostomatida Insecta Intramacronucleata Komagataella pastoris Komagataella Laurasiatheria Lepidoptera Liliopsida Magnoliopsida Mamestra brassicae Mamestra Mammalia Mandibulata Mesangiospermae Metazoa Muridae Murinae Muroidea Myomorpha Neognathae Neolepidoptera Neoptera Noctuidae Noctuoidea Obtectomera Oligohymenophorea Opisthokonta Ovis aries Ovis Panarthropoda Pancrustacea Pecora Pentapetalae Perdicinae Petrosaviidae Phaffomycetaceae Phasianidae Phasianinae Poales Primates Protostomia Pterygota RTA clade Rattus Rodentia Ruminantia Saccharomycetales Saccharomycetes Saccharomycotina Sar Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Spermatophyta Spodoptera frugiperda Spodoptera Streptophyta Streptophytina Suidae Suina Sus scrofa Sus Teleostomi Tetrahymena pyriformis Tetrahymena Tetrahymenidae Tetrahymenina Tetrapoda Theria Theropoda Tracheophyta Vertebrata Viridiplantae cellular organisms commelinids eudicotyledons malvids rosids saccharomyceta unclassified Anas unclassified entries unclassified sequences unidentified
G27243WR	Man(a1-6)Man(b1-4)GlcNAc(b1-4)GlcNAc(a1-		Linkage defined saccharide	748.27	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G37216UM	Man(a1-3)Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-		Linkage defined saccharide	748.27	50 kb inversion clade Amniota Amphiesmenoptera Amphipyrinae Archelosauria Archosauria Arthropoda Artiodactyla Ascomycota Aspergillaceae Aspergillus awamori Aspergillus Aves Bilateria Boreoeutheria Brassicales Carica papaya Carica Caricaceae Carnivora Chordata Coelurosauria Coturnix coturnix Coturnix japonica Coturnix Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dicondylia Dikarya Dinosauria Dipnotetrapodomorpha Ditrysia Ecdysozoa Embryophyta Endopterygota Euarchontoglires Eukaryota Eumetazoa Euphyllophyta Eurotiales Eurotiomycetes Eurotiomycetidae Euteleostomi Eutheria Fabaceae Fabales Felidae Feliformia Felinae Felis catus Felis Fungi Galliformes Galloanserae Glires Glossata Gnathostomata Gunneridae Heteroneura Hexapoda Insecta Komagataella pastoris Komagataella Laurasiatheria Lepidoptera Magnoliopsida Mammalia Mandibulata Mesangiospermae Metazoa Muroidea Myomorpha NPAAA clade Neognathae Neolepidoptera Neoptera Noctuidae Noctuoidea Obtectomera Opisthokonta Panarthropoda Pancrustacea Papilionoideae Pentapetalae Perdicinae Pezizomycotina Phaffomycetaceae Phaseoleae Phasianidae Protostomia Pterygota Rodentia Saccharomycetales Saccharomycetes Saccharomycotina Sarcopterygii Sauria Saurischia Sauropsida Spermatophyta Spodoptera frugiperda Spodoptera Streptophyta Streptophytina Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Theropoda Tracheophyta Vertebrata Vigna radiata Vigna Viridiplantae cellular organisms eudicotyledons fabids indigoferoid/millettioid clade leotiomyceta malvids rosids saccharomyceta unclassified entries unclassified sequences unidentified
G38582HO	Man(b1-4)GlcNAc(b1-4)Man(b1-4)GlcNAc(?1-		Linkage defined saccharide	748.27	Amniota Artiodactyla Bilateria Boreoeutheria Bovidae Capra hircus Capra Caprinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G64604UL	Gal(b1-4)GlcNAc(b1-3)Gal(b1-3)GalNAc(?1-	N-acetyllactosamine O-Glycan core 1 fuzzy Type 2 LN	Linkage defined saccharide	748.27	Amniota Artiodactyla Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Digenea Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Lophotrochozoa Mammalia Metazoa Opisthokonta Platyhelminthes Primates Protostomia Sarcopterygii Schistosoma mansoni Schistosoma Schistosomatidae Schistosomatoidea Simiiformes Spiralia Strigeidida Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Trematoda Vertebrata cellular organisms
G77627AJ	Gal(b1-3)[Gal(b1-4)GalNAc(b1-6)]GalNAc(?1-	O-Glycan core 1 fuzzy	Linkage defined saccharide	748.27	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G81133FA	Man(b1-4)GlcNAc(b1-4)Man(b1-4)GlcNAc		Linkage defined saccharide	748.27	Amniota Artiodactyla Bilateria Boreoeutheria Bovidae Capra hircus Capra Caprinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G96017QA	Gal(b1-3)[Gal(b1-4)GlcNAc(b1-6)]GalNAc	N-acetyllactosamine O-Glycan core 2 fuzzy Type 2 LN	Linkage defined saccharide	748.27	Amniota Artiodactyla Bilateria Boreoeutheria Bovidae Caprinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Opisthokonta Ovis aries Ovis Pecora Primates Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G01784XC	Fuc(a1-2)Gal(b1-3)GlcNAc(b1-3)GalNAc-ol	Type 1 LN	Linkage defined saccharide	734.3	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Pecora Primates Rattus Rodentia Ruminantia Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G02020YO	GalNAc(a1-3)[Fuc(a1-2)]Gal(b1-3)GalNAc-ol		Linkage defined saccharide	734.3	Amniota Amphibia Anura Artiodactyla Batrachia Bilateria Boreoeutheria Bufo bufo Bufo Bufonidae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Hyloidea Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Neobatrachia Opisthokonta Pipidae Pipoidea Primates Rattus Rodentia Sarcopterygii Silurana Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata Xenopodinae Xenopus tropicalis Xenopus cellular organisms
G15446MA	Fuc(a1-2)Gal(b1-3)[GalNAc(b1-6)]GalNAc-ol		Linkage defined saccharide	734.3	Amniota Artiodactyla Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Sarcopterygii Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G59354GZ	GalNAc(a1-3)[Fuc(a1-2)]Gal(b1-4)GalNAc-ol		Linkage defined saccharide	734.3	Amniota Artiodactyla Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Sarcopterygii Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G77674IK	Fuc(a1-2)Gal(b1-3)[GlcNAc(b1-6)]GalNAc-ol		Linkage defined saccharide	734.3	Amniota Amphibia Anura Artiodactyla Batrachia Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Lithobates palustris Lithobates Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Neobatrachia Opisthokonta Pecora Pipidae Pipoidea Primates Rana arvalis Rana temporaria Rana Rana Ranidae Ranoidea Rattus Rodentia Ruminantia Sarcopterygii Silurana Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata Xenopodinae Xenopus laevis Xenopus tropicalis Xenopus Xenopus cellular organisms
G81471KU	Fuc(a1-2)Gal(b1-4)GlcNAc(b1-3)GalNAc-ol	N-acetyllactosamine Type 2 LN	Linkage defined saccharide	734.3	Ambystoma mexicanum Ambystoma Ambystomatidae Amniota Amphibia Artiodactyla Batrachia Bilateria Boreoeutheria Catarrhini Caudata Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Primates Rattus Rodentia Salamandroidea Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms