GlyCosmos Portal

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GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	August 18, 2024

Displaying entries **151 - 175** of **1810** in total

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GlyTouCan ID ▼	IUPAC Condensed	Motifs	Subsumption Level	Monoisotopic Mass	Organisms
G89877QI	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan complex Polysialic acid	Linkage defined saccharide	2879.01	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Caprinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Lagomorpha Laurasiatheria Leporidae Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Oryctolagus cuniculus Oryctolagus Ovis aries Ovis Pecora Primates Rodentia Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G89864BN	Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan high mannose	Linkage defined saccharide	1720.59	Amniota Archelosauria Archosauria Artiodactyla Aves Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Coelurosauria Coturnix japonica Coturnix Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Neognathae Opisthokonta Pecora Perdicinae Phasianidae Primates Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms
G89663TB	GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-4)][GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	N-Glycan complex	Linkage defined saccharide	1665.62	Amniota Archelosauria Archosauria Artiodactyla Aves Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Neognathae Opisthokonta Pecora Phasianidae Phasianinae Primates Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms
G89659QV	Fuc(a1-2)Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-3)Gal(b1-4)Glc(b1-	Blood group H (type 2) Blood group H (type 2) Leb Lewis b Ley Paragloboside	Linkage defined saccharide	999.36	Amniota Bilateria Boreoeutheria Canidae Caniformia Canis lupus familiaris Canis lupus Canis Carnivora Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Opisthokonta Primates Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G89654YT	GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-2)[GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2030.76	Amniota Artiodactyla Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Sarcopterygii Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G89637WE	GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	N-Glycan complex	Linkage defined saccharide	1462.54	Acrididae Acrididea Acridoidea Acridomorpha Amniota Archelosauria Archosauria Arthropoda Artiodactyla Aves Batoidea Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Caelifera Catarrhini Chondrichthyes Chordata Coelurosauria Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dicondylia Dinosauria Dipnotetrapodomorpha Ecdysozoa Elasmobranchii Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Haplorrhini Hexapoda Hominidae Homininae Hominoidea Homo sapiens Homo Insecta Laurasiatheria Locusta migratoria Locusta Mammalia Mandibulata Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neognathae Neoptera Oedipodinae Opisthokonta Orthoptera Panarthropoda Pancrustacea Pecora Phasianidae Phasianinae Polyneoptera Primates Protostomia Pterygota Rodentia Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Tetronarce californica Tetronarce Theria Theropoda Torpedinidae Torpediniformes Vertebrata cellular organisms
G89544LF	Neu5Gc(a2-3)Gal(b1-3)GalNAc-ol		Linkage defined saccharide	692.25	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Lagomorpha Laurasiatheria Leporidae Mammalia Metazoa Opisthokonta Oryctolagus cuniculus Oryctolagus Pecora Ruminantia Sarcopterygii Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G89398MH	Fuc(a1-2)Gal(b1-3)[GalNAc(a1-3)[Fuc(a1-2)]Gal(b1-4)GlcNAc(b1-6)]GalNAc(?1-	Blood group A (type 2) O-Glycan core 2 fuzzy	Linkage defined saccharide	1243.47	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Primates Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G89398CR	GalNAc3/4S(b1-4)GlcNAc(b1-2)Man(a1-3)[Man(a1-3)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan hybrid	Linkage defined saccharide	1704.55	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G89285QP	GalNAc(b1-4)GlcNAc(b1-2)Man(a1-3)[GalNAc(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	LDN LacdiNAc N-Glycan complex	Linkage defined saccharide	1868.7	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Primates Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G89233QM	Gal(b1-3)Gal(b1-4)GlcNAc(b1-3)[Gal(b1-3)Gal(b1-4)GlcNAc(b1-6)]Gal(b1-4)GlcNAc(b1-3)Gal(b1-3)GalNAc-ol	I' antigen	Linkage defined saccharide	1804.66	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G89210OT	Gal(b1-3)[Gal(b1-3)Gal(b1-4)GlcNAc(b1-6)]GalNAc(a1-	N-acetyllactosamine O-Glycan core 2 Type 2 LN	Linkage defined saccharide	910.33	Amniota Artiodactyla Bilateria Boreoeutheria Bovidae Caprinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Ovis aries Ovis Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G89134YE	Neu5Ac(a2-3)Gal(b1-4)GlcNAc6S(b1-3)Gal	GM4	Linkage defined saccharide	916.25	Amniota Artiodactyla Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Sarcopterygii Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G88998CC	Glc(a1-2)Man(a1-2)Man(a1-2)Man(a1-3)[Glc(a1-2)Man(a1-3)[Glc(a1-2)Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan high mannose	Linkage defined saccharide	2206.75	Amniota Artiodactyla Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Sarcopterygii Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G88992QY	GalNAc?S(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1907.63	Amniota Artiodactyla Bilateria Boreoeutheria Bovidae Caprinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Ovis aries Ovis Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G88978UV	GalNAc(b1-3)GalNAc(?1-		Linkage defined saccharide	424.17	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G88904IL	Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Man(a1-2)Man(a1-3)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[D-Fuc(a1-6)]GlcNAc-ol	N-acetyllactosamine Type 2 LN	Linkage defined saccharide	1747.64	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G88876JQ	GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex	Linkage defined saccharide	1316.49	Acer pseudoplatanus Acer Acereae Actinopteri Actinopterygii Amniota Anas sp. Anas Anatidae Anatinae Anseriformes Archelosauria Archosauria Artiodactyla Aves Batoidea Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chondrichthyes Chordata Clupeocephala Coelurosauria Coturnix japonica Coturnix Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Dinosauria Dipnotetrapodomorpha Elasmobranchii Embryophyta Euarchontoglires Eukaryota Eumetazoa Euphyllophyta Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Gunneridae Haplorrhini Hippocastanoideae Hominidae Homininae Hominoidea Homo sapiens Homo Lagomorpha Laurasiatheria Leporidae Magnoliopsida Mammalia Mesangiospermae Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neognathae Neopterygii Opisthokonta Oryctolagus cuniculus Oryctolagus Ostariophysi Osteoglossocephalai Otomorpha Otophysi Pecora Pentapetalae Perdicinae Phasianidae Phasianinae Primates Rattus norvegicus Rattus Rodentia Ruminantia Sapindaceae Sapindales Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Spermatophyta Streptophyta Streptophytina Teleostei Teleostomi Tetrapoda Tetronarce californica Tetronarce Theria Theropoda Torpedinidae Torpediniformes Tracheophyta Vertebrata Viridiplantae cellular organisms eudicotyledons malvids rosids unclassified Anas unclassified entries unclassified sequences unidentified
G88864DI	Neu5Ac(a2-6)Gal(b1-4)GlcNAc(b1-2)[Gal(a1-3)Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Gal(a1-3)Gal(b1-4)GlcNAc(b1-2)[Gal(a1-3)Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN a-Gal antigen	Linkage defined saccharide	3294.17	Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Pecora Ruminantia Sarcopterygii Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G88800UV	Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)[Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2370.86	Amniota Artiodactyla Batoidea Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Carnivora Catarrhini Chondrichthyes Chordata Craniata Deuterostomia Dipnotetrapodomorpha Elasmobranchii Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Felidae Feliformia Felinae Felis catus Felis Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Pecora Primates Rodentia Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Tetronarce californica Tetronarce Theria Torpedinidae Torpediniformes Vertebrata cellular organisms
G88725FA	Neu5Ac(a2-3)Gal(b1-3)[Gal(b1-4)GlcNAc6S(b1-6)]GalNAc-ol	GM4	Linkage defined saccharide	1121.34	Amniota Archelosauria Archosauria Artiodactyla Aves Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Coelurosauria Craniata Deuterostomia Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Neognathae Opisthokonta Pecora Phasianidae Phasianinae Primates Ruminantia Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Teleostomi Tetrapoda Theria Theropoda Vertebrata cellular organisms
G88507CA	Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)Man(b1-4)GlcNAc(b1-4)GlcNAc		Linkage defined saccharide	1234.43	Amniota Artiodactyla Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Opisthokonta Sarcopterygii Suidae Suina Sus scrofa Sus Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G88495BI	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex Polysialic acid	Linkage defined saccharide	2222.78	Actinopteri Actinopterygii Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Clupeocephala Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Cypriniformes Cypriniphysae Cyprinoidei Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleosteomorpha Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Leuciscidae Mammalia Mesocricetus auratus Mesocricetus Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neopterygii Opisthokonta Osmeriformes Ostariophysi Osteoglossocephalai Otomorpha Otophysi Pecora Plecoglossidae Plecoglossus altivelis Plecoglossus Primates Pseudaspininae Pseudaspius hakonensis Pseudaspius Rodentia Ruminantia Sarcopterygii Simiiformes Stomiati Teleostei Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G88469YE	Man(a1-3)[Man(a1-6)]Man(b1-4)GlcNAc(?1-		Linkage defined saccharide	707.25	Alphavirus Alsuviricetes Amniota Archelosauria Archosauria Articulavirales Artiodactyla Ascomycota Aves Bilateria Boreoeutheria Bovidae Caprinae Carnivora Catarrhini Chordata Coelurosauria Coturnix coturnix Coturnix Craniata Deuterostomia Dikarya Dinosauria Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Felidae Feliformia Felinae Felis catus Felis Fungi Galliformes Galloanserae Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Insthoviricetes Kitrinoviricota Komagataella pastoris Komagataella Laurasiatheria Mammalia Martellivirales Metazoa Muridae Murinae Muroidea Myomorpha Negarnaviricota Neognathae Opisthokonta Orthomyxoviridae Orthornavirae Ovis aries Ovis Pecora Perdicinae Phaffomycetaceae Phasianidae Polyploviricotina Primates Rattus Riboviria Rodentia Ruminantia Saccharomycetales Saccharomycetes Saccharomycotina Sarcopterygii Sauria Saurischia Sauropsida Semliki Forest virus Simiiformes Teleostomi Tetrapoda Theria Theropoda Togaviridae Vertebrata Viruses cellular organisms saccharomyceta unclassified Orthomyxoviridae unclassified entries unclassified sequences unidentified influenza virus unidentified
G88437QR	Neu5Ac(a2-3)Gal(b1-3)GlcNAc	GM4	Linkage defined saccharide	674.24	Amniota Archelosauria Archosauria Aves Bifurcata Bilateria Boreoeutheria Calloselasma rhodostoma Calloselasma Catarrhini Chordata Coelurosauria Colubroidea Craniata Crotalinae Deuterostomia Dinosauria Dipnotetrapodomorpha Episquamata Equidae Equus caballus Equus Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Lepidosauria Mammalia Metazoa Opisthokonta Perissodactyla Primates Sarcopterygii Sauria Saurischia Sauropsida Serpentes Simiiformes Squamata Teleostomi Tetrapoda Theria Theropoda Toxicofera Unidentata Vertebrata Viperidae cellular organisms