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GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	August 18, 2024

Displaying entries **1 - 25** of **677** in total

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GlyTouCan ID	IUPAC Condensed ▼	Motifs	Subsumption Level	Monoisotopic Mass	Organisms
G29255IL	Neu5Ac(a2-?)Gal(b1-?)GlcNAc(b1-?)Man(a1-3)[Neu5Ac(a2-?)Gal(b1-?)GlcNAc(b1-?)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	N-Glycan complex Polysialic acid Type 1 LN	Linkage defined saccharide	2368.84	Amniota Betacoronavirus Bilateria Boreoeutheria Catarrhini Chordata Cornidovirineae Coronaviridae Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Nidovirales Opisthokonta Orthocoronavirinae Orthornavirae Pisoniviricetes Pisuviricota Primates Riboviria Rodentia Sarbecovirus Sarcopterygii Severe acute respiratory syndrome coronavirus 2 Severe acute respiratory syndrome-related coronavirus Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G55650WW	Neu5Ac(a2-?)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-?)Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-4)][Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan hybrid Polysialic acid	Linkage defined saccharide	3074.07	Artverviricota Human immunodeficiency virus 1 Lentivirus Ortervirales Orthoretrovirinae Pararnavirae Retroviridae Revtraviricetes Riboviria Viruses
G36280LM	Neu5Ac(a2-?)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-?)Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-4)][Man(a1-2)Man(a1-3)[Man(a1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan hybrid Polysialic acid	Linkage defined saccharide	3074.07	Artverviricota Human immunodeficiency virus Lentivirus Ortervirales Orthoretrovirinae Pararnavirae Primate lentivirus group Retroviridae Revtraviricetes Riboviria Viruses unclassified Primate lentivirus group
G24371XA	Neu5Ac(a2-?)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-?)Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-4)][Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan complex Polysialic acid	Linkage defined saccharide	2790.99	Artverviricota Human immunodeficiency virus 1 Lentivirus Ortervirales Orthoretrovirinae Pararnavirae Retroviridae Revtraviricetes Riboviria Viruses
G85501TK	Neu5Ac(a2-?)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-?)Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[GlcNAc(b1-4)][Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex Polysialic acid	Linkage defined saccharide	2790.99	Artverviricota Human immunodeficiency virus Lentivirus Ortervirales Orthoretrovirinae Pararnavirae Primate lentivirus group Retroviridae Revtraviricetes Riboviria Viruses unclassified Primate lentivirus group
G86750HK	Neu5Ac(a2-?)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-?)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	N-Glycan complex Polysialic acid	Linkage defined saccharide	2368.84	Amniota Artverviricota Bilateria Blubervirales Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hepadnaviridae Hepatitis B virus Hominidae Homininae Hominoidea Homo sapiens Homo Human immunodeficiency virus 1 Lentivirus Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Ortervirales Orthohepadnavirus Orthoretrovirinae Pararnavirae Primates Rattus norvegicus Rattus Retroviridae Revtraviricetes Riboviria Rodentia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G13346MU	Neu5Ac(a2-?)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-?)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan complex Polysialic acid	Linkage defined saccharide	2368.84	Artverviricota Blubervirales Hepadnaviridae Hepatitis B virus Human immunodeficiency virus Lentivirus Ortervirales Orthohepadnavirus Orthoretrovirinae Pararnavirae Primate lentivirus group Retroviridae Revtraviricetes Riboviria Viruses unclassified Primate lentivirus group
G53276NK	Neu5Ac(a2-?)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-?)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan complex Polysialic acid	Linkage defined saccharide	2222.78	Amniota Artverviricota Bilateria Blubervirales Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hepadnaviridae Hepatitis B virus Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Orthohepadnavirus Pararnavirae Primates Rattus norvegicus Rattus Revtraviricetes Riboviria Rodentia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms
G17652MS	Neu5Ac(a2-?)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-?)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex Polysialic acid	Linkage defined saccharide	2222.78	Artverviricota Blubervirales Hepadnaviridae Hepatitis B virus Orthohepadnavirus Pararnavirae Revtraviricetes Riboviria Viruses
G80573TM	Neu5Ac(a2-?)Gal(b1-4)GlcNAc	GM4	Linkage defined saccharide	674.24	Artverviricota Human immunodeficiency virus 1 Lentivirus Ortervirales Orthoretrovirinae Pararnavirae Retroviridae Revtraviricetes Riboviria Viruses
G36123IU	Neu5Ac(a2-6)GalNAc(a1-	Sialyl Tn antigen	Linkage defined saccharide	512.19	Amniota Artiodactyla Artverviricota Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Cestoda Chordata Craniata Cyclophyllidea Deuterostomia Digenea Dipnotetrapodomorpha Discoba Echinostomata Echinostomatoidea Euarchontoglires Eucestoda Euglenozoa Eukaryota Eumetazoa Euteleostomi Eutheria Fasciola hepatica Fasciola Fasciolidae Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Human immunodeficiency virus Kinetoplastea Lagomorpha Laurasiatheria Lentivirus Leporidae Lophotrochozoa Mammalia Mesocestoides vogae Mesocestoides Mesocestoididae Metakinetoplastina Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Ortervirales Orthoretrovirinae Oryctolagus cuniculus Oryctolagus Pararnavirae Pecora Plagiorchiida Platyhelminthes Primate lentivirus group Primates Protostomia Rattus norvegicus Rattus Retroviridae Revtraviricetes Riboviria Rodentia Ruminantia Sarcopterygii Schizotrypanum Simiiformes Spiralia Teleostomi Tetrapoda Theria Trematoda Trypanosoma cruzi Trypanosoma Trypanosomatida Trypanosomatidae Vertebrata Viruses cellular organisms unclassified Primate lentivirus group
G11346ER	Neu5Ac(a2-3/6)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3/6)Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Neu5Ac(a2-3/6)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3/6)Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc-ol	Polysialic acid	Linkage defined saccharide	3683.31	Amniota Artverviricota Bilateria Boreoeutheria Catarrhini Chordata Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Friend spleen focus-forming virus Gammaretrovirus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Ortervirales Orthoretrovirinae Pararnavirae Primates Rattus Retroviridae Revtraviricetes Riboviria Rodentia Sarcopterygii Simiiformes Spleen focus-forming virus Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms unclassified Gammaretrovirus
G35748HH	Neu5Ac(a2-3/6)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3/6)Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Neu5Ac(a2-3/6)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3/6)Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	N-Glycan complex Polysialic acid	Linkage defined saccharide	3681.3	Amniota Artverviricota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Friend spleen focus-forming virus Gammaretrovirus Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Ortervirales Orthoretrovirinae Pararnavirae Rattus Retroviridae Revtraviricetes Riboviria Rodentia Sarcopterygii Spleen focus-forming virus Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms unclassified Gammaretrovirus
G56039JB	Neu5Ac(a2-3/6)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-4)][Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan hybrid Polysialic acid	Linkage defined saccharide	2093.74	Artverviricota Friend murine leukemia virus Gammaretrovirus Murine leukemia virus Ortervirales Orthoretrovirinae Pararnavirae Retroviridae Revtraviricetes Riboviria Viruses
G51743KG	Neu5Ac(a2-3/6)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-2)Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	N-Glycan hybrid Polysialic acid	Linkage defined saccharide	2442.88	Amniota Artverviricota Bilateria Boreoeutheria Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Friend spleen focus-forming virus Gammaretrovirus Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Opisthokonta Ortervirales Orthoretrovirinae Pararnavirae Rattus Retroviridae Revtraviricetes Riboviria Rodentia Sarcopterygii Spleen focus-forming virus Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms unclassified Gammaretrovirus
G21459ST	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-4)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	N-Glycan complex Polysialic acid	Linkage defined saccharide	3025.07	Alpharhabdovirinae Haploviricotina Monjiviricetes Mononegavirales Negarnaviricota Orthornavirae Rhabdoviridae Riboviria Vesicular stomatitis virus Vesiculovirus Viruses unclassified Vesiculovirus
G49935CA	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-4)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan complex Polysialic acid	Linkage defined saccharide	3025.07	Alpharhabdovirinae Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Chordata Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Haploviricotina Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Monjiviricetes Mononegavirales Muroidea Myomorpha Negarnaviricota Opisthokonta Orthornavirae Pecora Primates Rhabdoviridae Riboviria Rodentia Ruminantia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata Vesicular stomatitis virus Vesiculovirus Viruses cellular organisms unclassified Vesiculovirus
G17689DH	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	N-Glycan complex Polysialic acid	Linkage defined saccharide	2368.84	Alphavirus Alsuviricetes Amniota Bilateria Boreoeutheria Catarrhini Cercopithecidae Cercopithecinae Cercopithecoidea Chlorocebus aethiops Chlorocebus Chordata Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Kitrinoviricota Mammalia Martellivirales Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Opisthokonta Orthornavirae Primates Rattus norvegicus Rattus Riboviria Rodentia Sarcopterygii Simiiformes Sindbis virus Teleostomi Tetrapoda Theria Togaviridae Vertebrata Viruses cellular organisms
G90889KG	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan complex Polysialic acid	Linkage defined saccharide	2368.84	Actinopteri Actinopterygii Alphavirus Alsuviricetes Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Catarrhini Cercopithecidae Cercopithecinae Cercopithecoidea Chlorocebus aethiops aethiops Chlorocebus aethiops Chlorocebus Chordata Clupeocephala Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Kitrinoviricota Laurasiatheria Mammalia Martellivirales Mesocricetus auratus Mesocricetus Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neopterygii Opisthokonta Orthornavirae Ostariophysi Osteoglossocephalai Otomorpha Otophysi Pecora Primates Rattus Riboviria Rodentia Ruminantia Sarcopterygii Simiiformes Sindbis virus Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Togaviridae Vertebrata Viruses cellular organisms unclassified entries unclassified sequences unidentified
G38955TH	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-4)][Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(b1-	N-Glycan hybrid Polysialic acid	Linkage defined saccharide	2239.8	Artverviricota Friend murine leukemia virus Gammaretrovirus Murine leukemia virus Ortervirales Orthoretrovirinae Pararnavirae Retroviridae Revtraviricetes Riboviria Viruses
G27571QK	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-4)][Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan hybrid Polysialic acid	Linkage defined saccharide	2239.8	Artverviricota Friend murine leukemia virus Gammaretrovirus Murine leukemia virus Ortervirales Orthoretrovirinae Pararnavirae Retroviridae Revtraviricetes Riboviria Viruses
G42686NP	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-4)][Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan hybrid Polysialic acid	Linkage defined saccharide	2093.74	Actinopteri Actinopterygii Artverviricota Bilateria Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Eukaryota Eumetazoa Euteleostomi Friend murine leukemia virus Gammaretrovirus Gnathostomata Metazoa Murine leukemia virus Neopterygii Opisthokonta Ortervirales Orthoretrovirinae Ostariophysi Osteoglossocephalai Otomorpha Otophysi Pararnavirae Retroviridae Revtraviricetes Riboviria Teleostei Teleostomi Vertebrata Viruses cellular organisms
G45560HM	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(b1-	N-Glycan complex Polysialic acid	Linkage defined saccharide	1931.69	Amniota Artverviricota Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Human immunodeficiency virus 1 Human immunodeficiency virus Lentivirus Mammalia Metazoa Opisthokonta Ortervirales Orthoretrovirinae Pararnavirae Primate lentivirus group Primates Retroviridae Revtraviricetes Riboviria Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms unclassified Primate lentivirus group
G68318SH	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex Polysialic acid	Linkage defined saccharide	1931.69	Amniota Artverviricota Bilateria Boreoeutheria Catarrhini Chordata Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Human immunodeficiency virus Lentivirus Mammalia Metazoa Muroidea Myomorpha Opisthokonta Ortervirales Orthoretrovirinae Pararnavirae Primate lentivirus group Primates Retroviridae Revtraviricetes Riboviria Rodentia Sarcopterygii Simiiformes Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms unclassified Primate lentivirus group
G01614ZM	Neu5Ac(a2-3)Gal(b1-3)[Neu5Ac(a2-6)]GalNAc(a1-	Di-sialyl T antigen	Linkage defined saccharide	965.33	Actinopteri Actinopterygii Amniota Artiodactyla Artverviricota Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Canidae Caniformia Canis lupus familiaris Canis lupus Canis Carnivora Catarrhini Chordata Clupeocephala Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Friend murine leukemia virus Gammaretrovirus Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Murine leukemia virus Muroidea Mus musculus Mus Mus Myomorpha Neopterygii Opisthokonta Ortervirales Orthoretrovirinae Ostariophysi Osteoglossocephalai Otomorpha Otophysi Pararnavirae Pecora Primates Rattus norvegicus Rattus Retroviridae Revtraviricetes Riboviria Rodentia Ruminantia Sarcopterygii Simiiformes Teleostei Teleostomi Tetrapoda Theria Vertebrata Viruses cellular organisms