GlyCosmos Portal

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GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	August 18, 2024

Displaying entries **326 - 350** of **393** in total

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GlyTouCan ID	IUPAC Condensed	Motifs	Subsumption Level	Monoisotopic Mass ▼	Organisms
G64973KT	Gal(b1-3)[Gal(b1-4)GlcNAc(b1-6)]GalNAc(a1-	N-acetyllactosamine O-Glycan core 2 Type 2 LN	Linkage defined saccharide	748.27	Actinopteri Actinopterygii Amniota Artiodactyla Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Caprinae Catarrhini Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Ovis aries Ovis Pecora Primates Rattus norvegicus Rattus Rodentia Ruminantia Sarcopterygii Simiiformes Teleostei Teleostomi Tetrapoda Theria Vertebrata artificial sequences cellular organisms other entries other sequences synthetic construct
G85237WB	GlcNAc(b1-3)[GalNAc(b1-4)]Gal(b1-4)Glc(?1-		Linkage defined saccharide	748.27	Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Eukaryota Eumetazoa Euteleostomi Gnathostomata Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Teleostei Teleostomi Vertebrata cellular organisms
G23006CO	GalNAc(b1-3)Gal(b1-4)Gal(b1-3)GalNAc(a1-	O-Glycan core 1 T antigen TF antigen	Linkage defined saccharide	748.27	Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Deuterostomia Eukaryota Eumetazoa Euteleosteomorpha Euteleostomi Gnathostomata Metazoa Neopterygii Oncorhynchus mykiss Oncorhynchus Opisthokonta Osteoglossocephalai Protacanthopterygii Salmonidae Salmoniformes Salmoninae Teleostei Teleostomi Vertebrata cellular organisms
G76881UY	GalNAc(b1-3)Gal(b1-4)Gal(b1-3)GalNAc(?1-	O-Glycan core 1 fuzzy	Linkage defined saccharide	748.27	Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Deuterostomia Eukaryota Eumetazoa Euteleosteomorpha Euteleostomi Gnathostomata Metazoa Neopterygii Oncorhynchus masou ishikawae Oncorhynchus masou Oncorhynchus mykiss Oncorhynchus nerka Oncorhynchus Opisthokonta Osteoglossocephalai Protacanthopterygii Salmonidae Salmoniformes Salmoninae Teleostei Teleostomi Vertebrata cellular organisms
G24508MA	Fuc(a1-3)GalNAc(b1-4)Gal(b1-3)GalNAc(a1-	O-Glycan core 1 T antigen TF antigen	Linkage defined saccharide	732.28	Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Eukaryota Eumetazoa Euteleostomi Gnathostomata Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Teleostei Teleostomi Vertebrata cellular organisms
G04762ZR	Kdn(a2-3)[Kdn(a2-6)]GalNAc-ol		Linkage defined saccharide	723.24	Actinopteri Actinopterygii Bilateria Botia Botiidae Chordata Clupeocephala Cobitidae Cobitinae Cobitoidei Craniata Cypriniformes Cypriniphysae Deuterostomia Eukaryota Eumetazoa Euteleostomi Gnathostomata Metazoa Misgurnus anguillicaudatus Misgurnus Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Teleostei Teleostomi Vertebrata cellular organisms
G72458LI	Kdn(a2-3)[Kdn(a2-6)]GalNAc(a1-	Tn antigen	Linkage defined saccharide	721.23	Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Cobitidae Cobitinae Cobitoidei Craniata Cypriniformes Cypriniphysae Deuterostomia Eukaryota Eumetazoa Euteleostomi Gnathostomata Metazoa Misgurnus anguillicaudatus Misgurnus Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Teleostei Teleostomi Vertebrata cellular organisms
G63334FZ	GlcNAc(b1-3)[Neu5Ac(a2-6)]GalNAc(a1-	O-Glycan core 3 Sialyl Tn antigen	Linkage defined saccharide	715.26	Actinopteri Actinopterygii Amniota Bilateria Boreoeutheria Catarrhini Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Primates Sarcopterygii Simiiformes Teleostei Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G48081MJ	GalNAc(a1-?)[Neu5Ac(a2-6)]GalNAc(?1-	O-Glycan core 5 fuzzy O-Glycan core 7 fuzzy	Linkage defined saccharide	715.26	Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Deuterostomia Eukaryota Eumetazoa Euteleosteomorpha Euteleostomi Gnathostomata Metazoa Neopterygii Oncorhynchus mykiss Oncorhynchus Opisthokonta Osteoglossocephalai Protacanthopterygii Salmonidae Salmoniformes Salmoninae Teleostei Teleostomi Vertebrata cellular organisms
G00407UJ	Gal(b1-3)GalNAc(b1-4)Gal(b1-4)Glc(?1-		Linkage defined saccharide	707.25	Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Eukaryota Eumetazoa Euteleostomi Gnathostomata Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Teleostei Teleostomi Vertebrata cellular organisms
G00061MO	GalNAc(b1-3)Gal(a1-4)Gal(b1-4)Glc(b1-	Gb4 Globoside P antigen	Linkage defined saccharide	707.25	Actinopteri Actinopterygii Amniota Archelosauria Archosauria Artiodactyla Autobranchia Aves Bilateria Bivalvia Boreoeutheria Bos taurus Bos Bovidae Bovinae Canidae Caniformia Canis lupus familiaris Canis lupus Canis Caprinae Carnivora Catarrhini Cercopithecidae Cercopithecinae Cercopithecoidea Chlorocebus aethiops Chlorocebus Chordata Clupeocephala Coelurosauria Craniata Cricetidae Cricetinae Deuterostomia Dinosauria Dipnotetrapodomorpha Discoba Euarchontoglires Euglenozoa Eukaryota Eumetazoa Euteleosteomorpha Euteleostomi Eutheria Galliformes Galloanserae Gallus gallus Gallus Glires Gnathostomata Gonideinae Haplorrhini Heteroconchia Hominidae Homininae Hominoidea Homo sapiens Homo Kinetoplastea Lagomorpha Laurasiatheria Leishmania amazonensis Leishmania mexicana species complex Leishmania Leishmania Leishmaniinae Leporidae Lophotrochozoa Mammalia Metakinetoplastina Metazoa Mollusca Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neognathae Neopterygii Oncorhynchus mykiss Oncorhynchus Opisthokonta Oryctolagus cuniculus Oryctolagus Osteoglossocephalai Ovis aries Ovis Palaeoheterodonta Pecora Phasianidae Phasianinae Primates Protacanthopterygii Protostomia Rattus Rodentia Ruminantia Salmonidae Salmoniformes Salmoninae Sarcopterygii Sauria Saurischia Sauropsida Simiiformes Sinohyriopsis schlegelii Sinohyriopsis Spiralia Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Theropoda Trypanosomatida Trypanosomatidae Unionida Unionidae Unionoidea Vertebrata cellular organisms
G11263YZ	Gal(b1-3)[Neu5Gc(a2-6)]GalNAc-ol		Linkage defined saccharide	692.25	Actinopteri Actinopterygii Amniota Amphibia Anura Artiodactyla Batrachia Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Bufo bufo Bufo Bufonidae Catarrhini Chordata Clupeocephala Craniata Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleosteomorpha Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Hyloidea Laurasiatheria Mammalia Metazoa Neobatrachia Neopterygii Oncorhynchus masou ishikawae Oncorhynchus masou Oncorhynchus mykiss Oncorhynchus Opisthokonta Osteoglossocephalai Pecora Primates Protacanthopterygii Ruminantia Salmonidae Salmoniformes Salmoninae Salvelinus leucomaenis japonicus Salvelinus leucomaenis pluvius Salvelinus leucomaenis Salvelinus Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G64527IJ	Neu5Gc(a2-3)Gal(b1-3)GalNAc(a1-	O-Glycan core 1 T antigen TF antigen	Linkage defined saccharide	690.23	Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Eukaryota Eumetazoa Euteleostomi Gnathostomata Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Teleostei Teleostomi Vertebrata cellular organisms
G64851ON	Gal(b1-3)[Neu5Gc(a2-6)]GalNAc(a1-	O-Glycan core 1 T antigen TF antigen	Linkage defined saccharide	690.23	Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Eukaryota Eumetazoa Euteleostomi Gnathostomata Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Teleostei Teleostomi Vertebrata cellular organisms
G49604IR	Gal(b1-3)[Neu5Gc(a2-6)]GalNAc(?1-	O-Glycan core 1 fuzzy	Linkage defined saccharide	690.23	Actinopteri Actinopterygii Amniota Artiodactyla Bilateria Boreoeutheria Chordata Clupeocephala Craniata Deuterostomia Dipnotetrapodomorpha Equidae Equus caballus Equus Eukaryota Eumetazoa Euteleosteomorpha Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Neopterygii Oncorhynchus mykiss Oncorhynchus Opisthokonta Osteoglossocephalai Perissodactyla Protacanthopterygii Salmonidae Salmoniformes Salmoninae Salvelinus leucomaenis pluvius Salvelinus leucomaenis Salvelinus Sarcopterygii Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G56682BC	Gal(b1-3)[Neu5Ac(a2-6)]GalNAc(a1-	O-Glycan core 1 Sialyl Tn antigen T antigen TF antigen	Linkage defined saccharide	674.24	Actinopteri Actinopterygii Amniota Artiodactyla Bilateria Boreoeutheria Canidae Caniformia Canis lupus familiaris Canis lupus Canis Carnivora Catarrhini Chordata Clupeocephala Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Primates Rattus norvegicus Rattus Rodentia Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Vertebrata artificial sequences cellular organisms other entries other sequences synthetic construct
G65562ZE	Neu5Ac(a2-3)Gal(b1-3)GalNAc(a1-	Sialyl T antigen	Linkage defined saccharide	674.24	Actinopteri Actinopterygii Amniota Artiodactyla Ascomycota Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Canidae Caniformia Canis lupus familiaris Canis lupus Canis Carnivora Catarrhini Chordata Clupeocephala Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Dikarya Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Fungi Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Laurasiatheria Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Pecora Primates Rattus norvegicus Rattus Rodentia Ruminantia Saccharomyces cerevisiae Saccharomyces Saccharomycetaceae Saccharomycetales Saccharomycetes Saccharomycotina Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Vertebrata cellular organisms saccharomyceta
G70444FA	Gal(b1-3)[Neu5Ac(a2-6)]GalNAc(?1-	O-Glycan core 1 fuzzy	Linkage defined saccharide	674.24	Actinopteri Actinopterygii Amniota Amphibia Anura Artiodactyla Batrachia Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Bufo japonicus Bufo Bufonidae Catarrhini Cercopithecidae Cercopithecinae Cercopithecoidea Chordata Clupeocephala Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dipnotetrapodomorpha Equidae Equus caballus Equus Euarchontoglires Eukaryota Eumetazoa Euteleosteomorpha Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Hyloidea Lagomorpha Laurasiatheria Leporidae Macaca radiata Macaca Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Neobatrachia Neopterygii Opisthokonta Oryctolagus cuniculus Oryctolagus Osteoglossocephalai Pecora Perissodactyla Primates Protacanthopterygii Rana dalmatina Rana Rana Ranidae Ranoidea Rattus norvegicus Rattus Rhinella arenarum Rhinella Rodentia Ruminantia Salmonidae Salmoniformes Salmoninae Salvelinus leucomaenis pluvius Salvelinus leucomaenis Salvelinus Sarcopterygii Simiiformes Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G81220GC	Fuc(a1-2)Gal(b1-4)[Fuc(a1-3)]Glc		Linkage defined saccharide	634.23	Actinopteri Actinopterygii Amniota Anguilliformes Bilateria Boreoeutheria Catarrhini Chordata Craniata Deuterostomia Dipnotetrapodomorpha Echidna Elopocephala Elopocephalai Elopomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Monotremata Muraenidae Muraeninae Neopterygii Opisthokonta Ornithorhynchidae Ornithorhynchus anatinus Ornithorhynchus Primates Prototheria Sarcopterygii Simiiformes Teleostei Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G09130SS	Gal(b1-3)[Kdn(a2-6)]GalNAc(a1-	O-Glycan core 1 T antigen TF antigen	Linkage defined saccharide	633.21	Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Eukaryota Eumetazoa Euteleostomi Gnathostomata Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Teleostei Teleostomi Vertebrata cellular organisms
G56516VH	Neu5Ac(a2-3)Gal(b1-4)Glc(?1-	GM4	Linkage defined saccharide	633.21	Actinopteri Actinopterygii Amniota Bilateria Boreoeutheria Catarrhini Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Primates Rodentia Sarcopterygii Simiiformes Teleostei Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G91237TK	Neu5Ac(a2-3)Gal(b1-4)Glc(b1-	GM3	Linkage defined saccharide	633.21	Acanthomorphata Actinopteri Actinopterygii African cichlids Amniota Amphibia Anura Archelosauria Archosauria Artiodactyla Aves Bacteria Batoidea Batrachia Bilateria Boreoeutheria Bos taurus Bos Bovidae Bovinae Campylobacter jejuni Campylobacter Campylobacteraceae Campylobacterales Campylobacterota Canidae Caniformia Canis lupus familiaris Canis lupus Canis Caprinae Carassius auratus Carassius Carnivora Catarrhini Cavia porcellus Cavia Caviidae Cercopithecidae Cercopithecinae Cercopithecoidea Channichthyidae Chinchilla Chinchillidae Chionodraco myersi Chionodraco Chondrichthyes Chordata Cichlidae Cichliformes Cichlomorphae Clupeocephala Coelurosauria Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Ctenosquamata Cyprinidae Cypriniformes Cyprininae Cypriniphysae Cyprinoidei Cyprinus carpio Cyprinus Deuterostomia Dinosauria Dipnotetrapodomorpha Discopyge tschudii Discopyge Elasmobranchii Enterobacterales Enterobacteriaceae Epsilonproteobacteria Escherichia coli Escherichia Euacanthomorphacea Euarchontoglires Eukaryota Eumetazoa Eupercaria Eurypterygia Euteleosteomorpha Euteleostomi Eutheria Felidae Feliformia Felinae Felis catus Felis Gadariae Gadidae Gadiformes Gadoidei Gadus morhua Gadus Galliformes Galloanserae Gallus gallus Gallus Gammaproteobacteria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Hystricomorpha Lagomorpha Laurasiatheria Leporidae Leuciscidae Leuciscinae Leucoraja erinacea Leucoraja Macaca mulatta Macaca Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Narcinidae Neognathae Neopterygii Neoteleostei Nototheniidae Notothenioidei Oncorhynchus mykiss Oncorhynchus Opisthokonta Oreochromini Oreochromis mossambicus Oreochromis Oryctolagus cuniculus Oryctolagus Ostariophysi Osteoglossocephalai Otomorpha Otophysi Ovalentaria Ovis aries Ovis Papio hamadryas Papio Paracanthopterygii Pecora Perciformes Percomorphaceae Phasianidae Phasianinae Phodopus sungorus Phodopus Pipidae Pipoidea Platyrrhini Primates Protacanthopterygii Pseudocrenilabrinae Pseudomonadota Rajidae Rajiformes Rattus Rodentia Ruminantia Salmonidae Salmoniformes Salmoninae Sarcopterygii Sauria Saurischia Sauropsida Scardinius erythrophthalmus Scardinius Simiiformes Suidae Suina Sus scrofa Sus Teleostei Teleostomi Tetrapoda Theria Theropoda Torpedinidae Torpediniformes Torpedo marmorata Torpedo Trematomus lepidorhinus Trematomus Vertebrata Xenopodinae Xenopus laevis Xenopus Xenopus Zeiogadaria cellular organisms
G31170TM	Kdn(a2-3)Gal(b1-4)Glc(b1-	Lactosylceramide	Linkage defined saccharide	592.19	Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Deuterostomia Eukaryota Eumetazoa Euteleosteomorpha Euteleostomi Gnathostomata Metazoa Neopterygii Oncorhynchus mykiss Oncorhynchus Opisthokonta Osteoglossocephalai Protacanthopterygii Salmonidae Salmoniformes Salmoninae Teleostei Teleostomi Vertebrata cellular organisms
G82699KR	Gal(b1-4)Gal(b1-3)GalNAc-ol		Linkage defined saccharide	547.21	Actinopteri Actinopterygii Amphibia Anura Batrachia Bilateria Chordata Clupeocephala Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleosteomorpha Euteleostomi Gnathostomata Metazoa Neobatrachia Neopterygii Opisthokonta Osteoglossocephalai Protacanthopterygii Rana temporaria Rana Rana Ranidae Ranoidea Salmonidae Salmoniformes Salmoninae Salvelinus leucomaenis japonicus Salvelinus leucomaenis pluvius Salvelinus leucomaenis Salvelinus Sarcopterygii Teleostei Teleostomi Tetrapoda Vertebrata cellular organisms
G04269FO	Gal?(?1-?)Gal?(?1-?)GalNAc(?1-		Glycosidic topology	545.2	Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Deuterostomia Eukaryota Eumetazoa Euteleosteomorpha Euteleostomi Gnathostomata Metazoa Neopterygii Oncorhynchus mykiss Oncorhynchus Opisthokonta Osteoglossocephalai Protacanthopterygii Salmonidae Salmoniformes Salmoninae Teleostei Teleostomi Vertebrata cellular organisms