GlyCosmos Portal

Table Filtering

GlyTouCan ID

IUPAC Condensed

Motifs

Subsumption Level

Monoisotopic Mass

Organisms

Taxonomy IDs

Tissues

Display
results per page.

Clear

Submissions

GlyCosmos Glycans

The list of validated glycans extracted from GlyTouCan, updated weekly.

Source	Last Updated
GlyTouCan	August 18, 2024

Displaying entries **276 - 300** of **393** in total

« First

‹ Prev

…

8

9

10

11

12

13

14

15

16

Next ›

Last »
GlyTouCan ID	IUPAC Condensed	Motifs	Subsumption Level	Monoisotopic Mass	Organisms ▼
G90913OZ	Kdn(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-4)][Kdn(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	N-Glycan complex N-acetyllactosamine Type 2 LN	Linkage defined saccharide	2489.87	Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Eukaryota Eumetazoa Euteleosteomorpha Euteleostomi Gnathostomata Metazoa Neopterygii Oncorhynchus mykiss Oncorhynchus Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Protacanthopterygii Salmonidae Salmoniformes Salmoninae Teleostei Teleostomi Vertebrata cellular organisms
G72458LI	Kdn(a2-3)[Kdn(a2-6)]GalNAc(a1-	Tn antigen	Linkage defined saccharide	721.23	Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Cobitidae Cobitinae Cobitoidei Craniata Cypriniformes Cypriniphysae Deuterostomia Eukaryota Eumetazoa Euteleostomi Gnathostomata Metazoa Misgurnus anguillicaudatus Misgurnus Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Teleostei Teleostomi Vertebrata cellular organisms
G78703NH	Kdn(a2-6)GalNAc(a1-	Tn antigen	Linkage defined saccharide	471.16	Actinopteri Actinopterygii Bilateria Chordata Clupeocephala Cobitidae Cobitinae Cobitoidei Craniata Cypriniformes Cypriniphysae Deuterostomia Eukaryota Eumetazoa Euteleostomi Gnathostomata Metazoa Misgurnus anguillicaudatus Misgurnus Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Teleostei Teleostomi Vertebrata cellular organisms
G04991IT	Kdn(a2-6)GalNAc-ol		Linkage defined saccharide	473.17	Actinopteri Actinopterygii Bilateria Botia Botiidae Chordata Clupeocephala Cobitoidei Craniata Cypriniformes Cypriniphysae Deuterostomia Eukaryota Eumetazoa Euteleostomi Gnathostomata Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Teleostei Teleostomi Vertebrata cellular organisms
G04762ZR	Kdn(a2-3)[Kdn(a2-6)]GalNAc-ol		Linkage defined saccharide	723.24	Actinopteri Actinopterygii Bilateria Botia Botiidae Chordata Clupeocephala Cobitidae Cobitinae Cobitoidei Craniata Cypriniformes Cypriniphysae Deuterostomia Eukaryota Eumetazoa Euteleostomi Gnathostomata Metazoa Misgurnus anguillicaudatus Misgurnus Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Teleostei Teleostomi Vertebrata cellular organisms
G48558GR	Neu5Ac(a2-3)[Gal(b1-3)GalNAc(b1-4)]Gal(b1-4)Glc(b1-	GM1	Linkage defined saccharide	998.34	Actinopteri Actinopterygii Bacteria Bilateria Campylobacter jejuni Campylobacter Campylobacteraceae Campylobacterales Campylobacterota Carassius auratus Carassius Chordata Clupeocephala Craniata Cyprinidae Cypriniformes Cyprininae Cypriniphysae Cyprinoidei Deuterostomia Discoba Epsilonproteobacteria Euglenozoa Eukaryota Eumetazoa Euteleostomi Gnathostomata Kinetoplastea Leishmania amazonensis Leishmania mexicana species complex Leishmania Leishmania Leishmaniinae Metakinetoplastina Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Teleostei Teleostomi Trypanosomatida Trypanosomatidae Vertebrata cellular organisms
G30588ZL					Actinopteri Actinopterygii Bacteria Betaproteobacteria Bilateria Chordata Clupeocephala Craniata Deuterostomia Enterobacterales Enterobacteriaceae Escherichia coli K1 Escherichia coli Escherichia Eukaryota Eumetazoa Euteleosteomorpha Euteleostomi Gammaproteobacteria Gnathostomata Mannheimia haemolytica Mannheimia Metazoa Moraxella nonliquefaciens Moraxella Moraxellaceae Moraxellales Neisseria meningitidis H44/76 Neisseria meningitidis serogroup B Neisseria meningitidis Neisseria Neisseriaceae Neisseriales Neopterygii Oncorhynchus mykiss Oncorhynchus nerka adonis Oncorhynchus nerka Oncorhynchus Opisthokonta Osteoglossocephalai Pasteurella Pasteurellaceae Pasteurellales Protacanthopterygii Pseudomonadota Salmo Salmonidae Salmoniformes Salmoninae Salvelinus namaycush Salvelinus Teleostei Teleostomi Vertebrata cellular organisms
G42686NP	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-4)][Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan hybrid Polysialic acid	Linkage defined saccharide	2093.74	Actinopteri Actinopterygii Artverviricota Bilateria Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Eukaryota Eumetazoa Euteleostomi Friend murine leukemia virus Gammaretrovirus Gnathostomata Metazoa Murine leukemia virus Neopterygii Opisthokonta Ortervirales Orthoretrovirinae Ostariophysi Osteoglossocephalai Otomorpha Otophysi Pararnavirae Retroviridae Revtraviricetes Riboviria Teleostei Teleostomi Vertebrata Viruses cellular organisms
G82699KR	Gal(b1-4)Gal(b1-3)GalNAc-ol		Linkage defined saccharide	547.21	Actinopteri Actinopterygii Amphibia Anura Batrachia Bilateria Chordata Clupeocephala Craniata Deuterostomia Dipnotetrapodomorpha Eukaryota Eumetazoa Euteleosteomorpha Euteleostomi Gnathostomata Metazoa Neobatrachia Neopterygii Opisthokonta Osteoglossocephalai Protacanthopterygii Rana temporaria Rana Rana Ranidae Ranoidea Salmonidae Salmoniformes Salmoninae Salvelinus leucomaenis japonicus Salvelinus leucomaenis pluvius Salvelinus leucomaenis Salvelinus Sarcopterygii Teleostei Teleostomi Tetrapoda Vertebrata cellular organisms
G60284TY	Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-4)][Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	Lewis x Lex N-Glycan hybrid N-acetyllactosamine SSEA-1 Type 2 LN	Linkage defined saccharide	1948.7	Actinopteri Actinopterygii Amniota Bilateria Boreoeutheria Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Mammalia Metazoa Muridae Murinae Muroidea Myomorpha Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Rattus norvegicus Rattus Rodentia Sarcopterygii Teleostei Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G53517XC	Neu5Gc(a2-3)Gal(?1-		Linkage defined saccharide	487.15	Actinopteri Actinopterygii Amniota Bilateria Boreoeutheria Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Dipnotetrapodomorpha Equidae Equus caballus Equus Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Laurasiatheria Mammalia Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Perissodactyla Sarcopterygii Teleostei Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G71502UE	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc	Polysialic acid	Linkage defined saccharide	2019.7	Actinopteri Actinopterygii Amniota Bilateria Boreoeutheria Catarrhini Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Leuciscidae Mammalia Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Primates Pseudaspininae Pseudaspius hakonensis Pseudaspius Sarcopterygii Simiiformes Teleostei Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G63160NR	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(?1-	Polysialic acid	Linkage defined saccharide	2019.7	Actinopteri Actinopterygii Amniota Bilateria Boreoeutheria Catarrhini Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Leuciscidae Mammalia Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Primates Pseudaspininae Pseudaspius hakonensis Pseudaspius Sarcopterygii Simiiformes Teleostei Teleostomi Tetrapoda Theria Vertebrata cellular organisms unclassified entries unclassified sequences unidentified
G29326OK	Neu5Ac(a2-8)Neu5Ac(a2-3)Gal(b1-3)[Neu5Ac(a2-8)Neu5Ac(a2-6)]GalNAc(a1-	Di-sialyl T antigen	Linkage defined saccharide	1547.52	Actinopteri Actinopterygii Amniota Bilateria Boreoeutheria Catarrhini Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Primates Sarcopterygii Simiiformes Teleostei Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G63334FZ	GlcNAc(b1-3)[Neu5Ac(a2-6)]GalNAc(a1-	O-Glycan core 3 Sialyl Tn antigen	Linkage defined saccharide	715.26	Actinopteri Actinopterygii Amniota Bilateria Boreoeutheria Catarrhini Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Primates Sarcopterygii Simiiformes Teleostei Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G42797SX	Neu5Ac(a2-3)[GalNAc(b1-4)]Gal(b1-3)GalNAc(a1-	Sialyl T antigen	Linkage defined saccharide	877.32	Actinopteri Actinopterygii Amniota Bilateria Boreoeutheria Catarrhini Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Primates Sarcopterygii Simiiformes Teleostei Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G49299ZV	Gal(b1-4)[Fuc(a1-3)]GlcNAc(b1-2)Man(a1-3)[GlcNAc(b1-4)][Man(a1-3)[Man(a1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)[Fuc(a1-6)]GlcNAc(?1-	Lewis x Lex N-Glycan hybrid N-acetyllactosamine SSEA-1 Type 2 LN	Linkage defined saccharide	2094.76	Actinopteri Actinopterygii Amniota Bilateria Boreoeutheria Catarrhini Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Primates Sarcopterygii Simiiformes Teleostei Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G74621DY	Gal(b1-4)Glc(?1-		Linkage defined saccharide	342.12	Actinopteri Actinopterygii Amniota Bilateria Boreoeutheria Catarrhini Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Primates Sarcopterygii Simiiformes Teleostei Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G56516VH	Neu5Ac(a2-3)Gal(b1-4)Glc(?1-	GM4	Linkage defined saccharide	633.21	Actinopteri Actinopterygii Amniota Bilateria Boreoeutheria Catarrhini Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Primates Rodentia Sarcopterygii Simiiformes Teleostei Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G46748BU	Neu5Ac(a2-3)Gal(b1-3)[Gal(b1-4)GlcNAc(b1-6)]GalNAc(a1-	N-acetyllactosamine O-Glycan core 2 Sialyl T antigen Type 2 LN	Linkage defined saccharide	1039.37	Actinopteri Actinopterygii Amniota Bilateria Boreoeutheria Catarrhini Chordata Clupeocephala Craniata Cypriniformes Cypriniphysae Cyprinoidei Danio rerio Danio Danionidae Danioninae Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muridae Murinae Muroidea Mus musculus Mus Mus Myomorpha Neopterygii Opisthokonta Ostariophysi Osteoglossocephalai Otomorpha Otophysi Primates Rattus norvegicus Rattus Rodentia Sarcopterygii Simiiformes Teleostei Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G42380GI	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan complex Polysialic acid	Linkage defined saccharide	3535.24	Actinopteri Actinopterygii Amniota Bilateria Boreoeutheria Catarrhini Chordata Clupeocephala Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleosteomorpha Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muroidea Myomorpha Neopterygii Opisthokonta Osteoglossocephalai Primates Protacanthopterygii Rodentia Salmonidae Salmoniformes Sarcopterygii Simiiformes Teleostei Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G17441MI	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex Polysialic acid	Linkage defined saccharide	2879.01	Actinopteri Actinopterygii Amniota Bilateria Boreoeutheria Catarrhini Chordata Clupeocephala Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleosteomorpha Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muroidea Myomorpha Neopterygii Opisthokonta Osmeriformes Osteoglossocephalai Plecoglossidae Plecoglossus altivelis Plecoglossus Primates Rodentia Sarcopterygii Simiiformes Stomiati Teleostei Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G56963QZ	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc(?1-	N-Glycan complex Polysialic acid	Linkage defined saccharide	3535.24	Actinopteri Actinopterygii Amniota Bilateria Boreoeutheria Catarrhini Chordata Clupeocephala Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleosteomorpha Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muroidea Myomorpha Neopterygii Opisthokonta Osmeriformes Osteoglossocephalai Plecoglossidae Plecoglossus altivelis Plecoglossus Primates Rodentia Sarcopterygii Simiiformes Stomiati Teleostei Teleostomi Tetrapoda Theria Vertebrata cellular organisms unclassified entries unclassified sequences unidentified
G42600SB	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-6)]Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan complex Polysialic acid	Linkage defined saccharide	2879.01	Actinopteri Actinopterygii Amniota Bilateria Boreoeutheria Catarrhini Chordata Clupeocephala Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleosteomorpha Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muroidea Myomorpha Neopterygii Opisthokonta Osmeriformes Osteoglossocephalai Plecoglossidae Plecoglossus altivelis Plecoglossus Primates Protacanthopterygii Rodentia Salmonidae Salmoniformes Sarcopterygii Simiiformes Stomiati Teleostei Teleostomi Tetrapoda Theria Vertebrata cellular organisms
G97709WW	Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-4)]Man(a1-3)[Neu5Ac(a2-3)Gal(b1-4)GlcNAc(b1-2)Man(a1-6)]Man(b1-4)GlcNAc(b1-4)GlcNAc	N-Glycan complex Polysialic acid	Linkage defined saccharide	2879.01	Actinopteri Actinopterygii Amniota Bilateria Boreoeutheria Catarrhini Chordata Clupeocephala Craniata Cricetidae Cricetinae Cricetulus griseus Cricetulus Deuterostomia Dipnotetrapodomorpha Euarchontoglires Eukaryota Eumetazoa Euteleosteomorpha Euteleostomi Eutheria Glires Gnathostomata Haplorrhini Hominidae Homininae Hominoidea Homo sapiens Homo Mammalia Metazoa Muroidea Myomorpha Neopterygii Opisthokonta Osmeriformes Osteoglossocephalai Plecoglossidae Plecoglossus altivelis Plecoglossus Primates Protacanthopterygii Rodentia Salmonidae Salmoniformes Sarcopterygii Simiiformes Stomiati Teleostei Teleostomi Tetrapoda Theria Vertebrata cellular organisms